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Many microswimmers are able to swim through viscous fluids by employing periodic non-reciprocal deformations of their appendages. Here we use a simple microswimmer model inspired by swimming biflagellates which consists of a spherical cell…

Soft Condensed Matter · Physics 2025-08-22 Sridhar Bulusu , Andreas Zöttl

In this article we study synchronization of systems of homogeneous phase-coupled oscillators with plastic coupling strengths and arbitrary underlying topology. The dynamics of the coupling strength between two oscillators is governed by the…

Dynamical Systems · Mathematics 2016-02-24 Andrey Gushchin , Enrique Mallada , Ao Tang

Hydrodynamic equations for an isotropic solution of active polar filaments are derived from a microscopic mean-field model of the forces exchanged between motors and filaments. We find that a spatial dependence of the motor stepping rate…

Soft Condensed Matter · Physics 2015-06-24 T. B. Liverpool , M. Cristina Marchetti

Flow-Induced Self-Assembly (FISA) is the phenomena of particle chaining in viscoelastic fluids while experiencing shear flow. FISA has a large number of applications across many fields including material science, food processing and…

Some microorganisms, such as spermatozoa, synchronize their flagella when swimming in close proximity. Using a simplified model (two infinite, parallel, two-dimensional waving sheets), we show that phase-locking arises from hydrodynamics…

Biological Physics · Physics 2009-09-15 Gwynn J. Elfring , Eric Lauga

The twisting and writhing of a cell body and associated mechanical stresses is an underappreciated constraint on microbial self-propulsion. Multi-flagellated bacteria can even buckle and writhe under their own activity as they swim through…

Soft Condensed Matter · Physics 2023-09-25 Wilson Lough , Douglas B. Weibel , Saverio E. Spagnolie

By taking into account the hydrodynamic interactions in a one dimensional array of model cilia attached to a no-slip cylinderical surface, we investigate their synchronized motion. We show, how does the emergence of metachronal waves depend…

Biological Physics · Physics 2017-05-31 Aref Ghorbani , Ali Najafi

The dynamics of filaments in flow are central to understanding a wide range of biological and soft-matter systems, yet their behavior under time-dependent forcing remains poorly understood. Here, we investigate the long-time dynamics of…

Soft Condensed Matter · Physics 2026-01-05 Francesco Bonacci , Brato Chakrabarti , Olivia du Roure , Anke Lindner , David Saintillan

Colloidal particles moving in a fluid interact via the induced velocity field. The collective dynamic state for a class of actively forced colloids, driven by harmonic potentials via a rule that couples forces to configurations, to perform…

Soft Condensed Matter · Physics 2011-10-11 Loïc Damet , Giovanni M. Cicuta , Jurij Kotar , Marco Cosentino Lagomarsino , Pietro Cicuta

Essentially all biology is active and dynamic. Biological entities autonomously sense, com- pute, and respond using energy-coupled ratchets that can produce force and do work. The cytoskeleton, along with its associated proteins and motors,…

Soft Condensed Matter · Physics 2017-03-28 Kasimira T. Stanhope , Vikrant Yadav , Christian D. Santangelo , Jennifer L. Ross

During the life of animals, epithelial tissues undergo extensive deformations--first to form organs during embryogensis and later to preserve integrity and function in adulthood. To what extent these deformations resemble that of non-living…

Soft Condensed Matter · Physics 2025-04-23 Urska Andrensek , Matej Krajnc

Cell crawling requires the generation of intracellular forces by the cytoskeleton and their transmission to an extracellular substrate through specific adhesion molecules. Crawling cells show many features of excitable systems, such as…

Biological Physics · Physics 2020-06-02 Pierre Sens

We investigate the transport of elastic active filaments in two-dimensional turbulence, focusing on how propulsion geometry and elasticity determine vortex trapping and transport. Using a bead-spring model with activity applied at the…

Fluid Dynamics · Physics 2026-05-26 Kunal Kumar , Aliv Sahoo , Rahul Kumar Singh , Samriddhi Sankar Ray

We study the segregation of motile semiflexible filaments from a background of similar but non-motile filaments. Our Langevin dynamics simulations reveal a wide range of emergent structures governed by filament flexibility and activity,…

Soft Condensed Matter · Physics 2025-12-22 Chitrak Bhowmik , Aparna Baskaran , Sriram Ramaswamy

Multi-flagellated bacteria utilize the hydrodynamic interaction between their filamentary tails, known as flagella, to swim and change their swimming direction in low Reynolds number flow. This interaction, referred to as bundling and…

Robotics · Computer Science 2023-07-04 Zhuonan Hao , Sangmin Lim , M. Khalid Jawed

A model of an autonomous three-sphere microswimmer is proposed by implementing a coupling effect between the two natural lengths of an elastic microswimmer. Such a coupling mechanism is motivated by the previous models for synchronization…

Soft Condensed Matter · Physics 2021-05-26 Katsutomo Era , Yuki Koyano , Yuto Hosaka , Kento Yasuda , Hiroyuki Kitahata , Shigeyuki Komura

Run-and-tumble (RT) motion is commonly observed in flagellated microswimmers, arising from synchronous and asynchronous flagellar beating. One such example is a biflagellated alga, called \textit{Chlamydomonas reinhardtii}. Its flagellar…

Soft Condensed Matter · Physics 2025-10-17 Somnath Paramanick , Umashankar Pardhi , Harsh Soni , Nitin Kumar

Micro-organisms propel themselves in viscous environments by the periodic, nonreciprocal beating of slender appendages known as flagella. Active materials have been widely exploited to mimic this form of locomotion. However, the realization…

Soft Condensed Matter · Physics 2024-08-06 Ariel Surya Boiardi , Giovanni Noselli

The bending of cilia and flagella is driven by forces generated by dynein motor proteins. These forces slide adjacent microtubule doublets within the axoneme, the motile cytoskeletal structure. To create regular, oscilla- tory beating…

Biological Physics · Physics 2015-06-19 V. Mukundan , P. Sartori , V. F. Geyer , F. Julicher , J. Howard

Viscoelastic flows through microscale porous arrays exhibit complex path-selection and switching phenomena. However, understanding this process is limited by a lack of studies linking between a single object and large arrays. Here, we…

Fluid Dynamics · Physics 2021-02-10 Cameron C. Hopkins , Simon J. Haward , Amy Q. Shen
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