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We report on the dynamics of collective alignment in groups of the cichlid fish, Etroplus suratensis. Focusing on small-to-intermediate sized groups ($10<N<100$), we demonstrate that schooling (highly polarised and coherent motion) is…

Statistical Mechanics · Physics 2020-04-22 Jitesh Jhawar , Richard G. Morris , U. R. Amith-Kumar , M. Danny Raj , Harikrishnan R. , Vishwesha Guttal

Fish schools are able to display a rich variety of collective states and behavioural responses when they are confronted to threats. However a school's response to perturbations may be different depending on its collective state. Here we use…

Quantitative Methods · Quantitative Biology 2015-02-04 Daniel S. Calovi , Ugo Lopez , Paul Schuhmacher , Hugues Chaté , Clément Sire , Guy Theraulaz

Fish schooling is often modeled with self-propelled particles subject to phenomenological behavioral rules. Although fish are known to sense and exploit flow features, these models usually neglect hydrodynamics. Here, we propose a novel…

Biological Physics · Physics 2018-05-16 Audrey Filella , François Nadal , Clément Sire , Eva Kanso , Christophe Eloy

A novel model of the wake of swimming fish is developed and incorporated into a dynamical model of a fish school to explore the effect of hydrodynamics on the emergent behavior in schooling fish. The model incorporates well-established…

Fluid Dynamics · Physics 2025-07-11 Ji Zhou , Jung-Hee Seo , Rajat Mittal

The complex formations exhibited by schooling fish have long been the object of fascination for biologists and physicists. However, the physical and sensory mechanisms leading to organized collective behavior remain elusive. On the physical…

Fluid Dynamics · Physics 2025-07-09 Anand U. Oza , Eva Kanso , Michael J. Shelley

There is increasing evidence that fish gain energetic benefits when they swim in a school. The most recent indications of such benefits are a lower tail (or fin) beat at the back of a school and reduced oxygen consumption in schooling fish…

Quantitative Methods · Quantitative Biology 2013-07-30 C. K. Hemelrijk , D. A. P. Reid , H. Hildenbrandt , J. T. Padding

While a rich variety of self-propelled particle models propose to explain the collective motion of fish and other animals, rigorous statistical comparison between models and data remains a challenge. Plausible models should be flexible…

Quantitative Methods · Quantitative Biology 2017-04-28 Maksym Romenskyy , James E. Herbert-Read , Ashley J. W. Ward , David J. T. Sumpter

Three-dimensional simulations with fully resolved hydrodynamics are performed to study the collective motion of model swimmers in confinement. We show that certain swimming mechanisms can lead to traveling wave-like collective motion even…

Soft Condensed Matter · Physics 2016-04-20 Norihiro Oyama , John Jairo Molina , Ryoichi Yamamoto

Schooling fish often self-organize into a variety of collective patterns, from polarized schooling to rotational milling. Mathematical models support the emergence of these large-scale patterns from local decentralized interactions, in the…

Adaptation and Self-Organizing Systems · Physics 2025-07-23 Alyssa Chan , Eva Kanso

Gas bubbles in a sound field are submitted to a radiative force, known as the secondary Bjerknes force. We propose an original experimental setup that allows us to investigate in details this force between two bubbles, as a function of the…

Soft Condensed Matter · Physics 2015-12-09 Maxime Lanoy , Caroline Derec , Arnaud Tourin , Valentin Leroy

In a companion paper, a reduced model for propagation of acoustic waves in a cloud of inertial cavitation bubbles was proposed. The wave attenuation was calculated directly from the energy dissipated by a single bubble, the latter being…

Fluid Dynamics · Physics 2013-02-26 Olivier Louisnard

Schooling, an archetype of collective behavior, emerges from the interactions of fish responding to visual and other informative cues mediated by their aqueous environment. In this context, a fundamental and largely unexplored question…

Crowd movements are observed among different species and on different scales, from insects to mammals, as well as in non-cognitive systems, such as motile cells. When forced to escape through a narrow opening, most terrestrial animals…

Biological Physics · Physics 2024-02-16 R. Larrieu , P. Moreau , C. Graff , P. Peyla , A. Dupont

We report experimental observations of the volume acoustic modes of air bubbles in water, including both the fundamental Minnaert breathing mode and a family of higher-order modes extending into the megahertz frequency range. Bubbles were…

Fluid Dynamics · Physics 2024-05-03 K. G. Scheuer , F. B. Romero , R. G. DeCorby

Bubbles display astonishing acoustical properties since they are able to absorb and scatter large amounts of energy coming from waves whose wavelengths are two orders of magnitude larger than the bubble size. Thus, as the interaction…

Fluid Dynamics · Physics 2012-11-07 Hector Estrada , Francisco Meseguer

Sound can exert forces on objects of any material and shape. This has made the contactless manipulation of objects by intense ultrasound a fascinating area of research with wide-ranging applications. While much is understood for acoustic…

Soft Condensed Matter · Physics 2024-01-17 Melody X. Lim , Bryan VanSaders , Heinrich M. Jaeger

One of the most highly debated questions in the field of animal swarming and social behaviour, is the collective random patterns and chaotic behaviour formed by some animal species, in particular if there is a danger. Is such a behaviour…

Populations and Evolution · Quantitative Biology 2017-01-04 Usama Kadri , Franz Brümmer , Anan Kadri

Fish in schooling formations navigate complex flow-fields replete with mechanical energy in the vortex wakes of their companions. Their schooling behaviour has been associated with evolutionary advantages including collective energy…

Fluid Dynamics · Physics 2022-06-08 Siddhartha Verma , Guido Novati , Petros Koumoutsakos

Micro-swimmers can serve as cargo carriers that move deep inside complex flow networks. When a school collectively entrains the surrounding fluid, their transport capacity can be enhanced. This effect is quantified with good agreement…

Soft Condensed Matter · Physics 2021-08-26 Chenyu Jin , Yibo Chen , Corinna C. Maass , Arnold J. T. M. Mathijssen

Collective behaviors displayed by groups of social animals are observed frequently in nature. Understanding and predicting the behavior of complex biological systems is dependent on developing effective descriptions and models. While…

Biological Physics · Physics 2020-07-01 Julia A. Giannini , James G. Puckett
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