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The habitat of microorganisms is typically complex and viscoelastic. A natural question arises: Do polymers in a suspension of active swimmers enhance chaotic motion or promote orientational order? We address this issue by performing…

Soft Condensed Matter · Physics 2025-12-23 Yuan Zhou , Qingzhi Zou , Ignacio Pagonabarraga , Kaihuan Zhang , Kai Qi

We run numerical simulations of strongly confined suspensions of model spherical swimmers called "squirmers". Because of the confinement, the Stokeslet dipoles generated by the particles are quickly screened and the far-field flow is…

Fluid Dynamics · Physics 2016-06-22 Jean-Baptiste Delfau , John Molina , Masaki Sano

Microswimmers in suspension exhibit collective swimming behaviour, forming various self-organised structures including ordered, aggregated, and turbulent-like structures. When mixed with passive particles phase-separation is known to occur,…

Fluid Dynamics · Physics 2026-03-26 Alexander Chamolly , Takuji Ishikawa

We use numerical simulations to probe the dynamics of concentrated suspensions of spherical microswimmers interacting hydrodynamically. Previous work in the dilute limit predicted orientational instabilities of aligned suspensions for both…

Soft Condensed Matter · Physics 2013-02-12 Arthur A. Evans , Takuji Ishikawa , Takami Yamaguchi , Eric Lauga

We present a new development of the force-coupling method (FCM) to address the accurate simulation of a large number of interacting micro-swimmers. Our approach is based on the squirmer model, which we adapt to the FCM framework, resulting…

Soft Condensed Matter · Physics 2015-12-21 Blaise Delmotte , Eric Keaveny , Franck Plouraboue , Eric Climent

The incessant swimming motion of microbes in dense suspensions can give rise to striking collective motions and coherent structures. However, theoretical investigations of these structures typically utilize either computationally demanding…

Fluid Dynamics · Physics 2019-05-29 Douglas R. Brumley , Timothy J. Pedley

The sedimentation process in a suspension of bacteria is the result of the competition between gravity and the intrinsic motion of the microorganisms. We perform simulations of run-and-tumble "squirmers" that move in a fluid medium,…

Soft Condensed Matter · Physics 2021-11-11 Andrea Scagliarini , Ignacio Pagonabarraga

Active particles, which interact hydrodynamically, display a remarkable variety of emergent collective phenomena. We use squirmers to model spherical microswimmers and explore the collective behavior of thousands of them under the influence…

Fluid Dynamics · Physics 2017-10-31 Jan-Timm Kuhr , Johannes Blaschke , Felix Rühle , Holger Stark

Active suspensions encompass a wide range of complex fluids containing microscale energy-injecting particles, such as cells, bacteria or artificially powered active colloids. Because they are intrinsically non-equilibrium, active…

Soft Condensed Matter · Physics 2024-07-09 Zhouyang Ge , Gwynn J. Elfring

We study the role of hydrodynamic interactions in the collective behaviour of collections of microscopic active particles suspended in a fluid. We introduce a novel calculational framework that allows us to separate the different…

Soft Condensed Matter · Physics 2017-09-06 Natsuhiko Yoshinaga , Tanniemola B. Liverpool

Collective behaviour in suspensions of microswimmers is often dominated by the impact of long-ranged hydrodynamic interactions. These phenomena include active turbulence, where suspensions of pusher bacteria at sufficient densities exhibit…

Soft Condensed Matter · Physics 2019-08-27 Dora Bardfalvy , Henrik Nordanger , Cesare Nardini , Alexander Morozov , Joakim Stenhammar

We study numerically the hydrodynamics of a self-propelled particle system, consisting of spherical squirmers sedimented on a flat surface. We observe the emergence of dynamic structures, due to the interplay of particle-particle and…

Soft Condensed Matter · Physics 2019-12-20 Zaiyi Shen , Juho S. Lintuvuori

Hydrodynamical interactions of active micro-particles are pervasive in our planet's fluid environments. Hence, understanding the interactions of these self-propelled particles is essential for science and engineering. In this paper the…

Soft Condensed Matter · Physics 2022-12-28 Yonatan Ashenafi

We study how hydrodynamic interactions affect the collective behaviour of active particles suspended in a fluid at high concentrations, with particular attention to lubrication forces which appear when the particles are very close to one…

Soft Condensed Matter · Physics 2018-05-29 Natsuhiko Yoshinaga , Tanniemola B. Liverpool

We present a two dimensional model of hydrodynamic interaction between a circular swimmer and a circular post at low Reynolds number, using a point singularity description of the swimming activity. We derive a nonlinear dynamical system…

Fluid Dynamics · Physics 2016-12-09 Dario Papavassiliou , Gareth P Alexander

We study the energy expenditure and structural correlations in semi-dilute to concentrated suspensions of squirmers using active fast Stokesian dynamics simulations. Specifically, we simulate apolar active suspensions of squirmers, or…

Soft Condensed Matter · Physics 2026-03-03 Zhouyang Ge , Gwynn J. Elfring

Self-propelled particles with hydrodynamic interactions (microswimmers) have previously been shown to produce long-range ordering phenomena. Many theoretical explanations for these collective phenomena are connected to instabilities in the…

Soft Condensed Matter · Physics 2017-05-03 Yuzhou Qian , Peter R. Kramer , Patrick T. Underhill

Microorganisms thrive in complex environments and their behavior in fluids holds significant importance for various medical and industrial applications. By conducting Lattice Boltzmann simulations, the transport and rotational properties of…

Soft Condensed Matter · Physics 2025-03-25 Yuan Zhou , Kai Qi , Marco De Corato , Kevin Stratford , Ignacio Pagonabarraga

The dynamic interactions between pairs of swimming microorganisms underpin the collective behaviour of larger suspensions, but accurately calculating pairwise collisions has typically required the use of numerical simulations in which…

Soft Condensed Matter · Physics 2021-11-30 C. Darveniza , T. Ishikawa , T. J. Pedley , D. R. Brumley

Previous particle-based computer simulations have revealed a significantly more pronounced tendency of spontaneous global polar ordering in puller (contractile) microswimmer suspensions than in pusher (extensile) suspensions. We here…

Soft Condensed Matter · Physics 2018-10-23 Christian Hoell , Hartmut Löwen , Andreas M. Menzel
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