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We present a comprehensive theory of the dynamics and fluctuations of a two-dimensional suspension of polar active particles in an incompressible fluid confined to a substrate. We show that, depending on the sign of a single parameter, a…

Soft Condensed Matter · Physics 2020-01-22 Ananyo Maitra , Pragya Srivastava , M. Cristina Marchetti , Sriram Ramaswamy , Martin Lenz

In order to asses the effect of hydrodynamics in the assembly of active attractive spheres, we simulate a semi-dilute suspension of attractive self-propelled spherical particles in a quasi two dimensional geometry comparing the case with…

Soft Condensed Matter · Physics 2016-12-15 Francisco Alarcón , Chantal Valeriani , Ignacio Pagonabarraga

We study the collective motion of confined spherical microswimmers such as active colloids which we model by so-called squirmers. To simulate hydrodynamic flow fields including thermal noise, we use the method of multi-particle collision…

Soft Condensed Matter · Physics 2014-03-20 Andreas Zöttl , Holger Stark

Microswimmers exhibit an intriguing, highly-dynamic collective motion with large-scale swirling and streaming patterns, denoted as active turbulence -- reminiscent of classical high-Reynolds-number hydrodynamic turbulence. Various…

Soft Condensed Matter · Physics 2024-06-03 Kai Qi , Elmar Westphal , Gerhard Gompper , Roland G. Winkler

We investigate how shear influences the dynamics of active particles in dilute to concentrated suspensions. Using apolar active suspensions of squirmers as model systems, we show how their long-time diffusive dynamics can surprisingly slow…

Soft Condensed Matter · Physics 2025-11-14 Zhouyang Ge , John F. Brady , Gwynn J. Elfring

We present the results of hydrodynamic simulations using the method of multi-particle collision dynamics for a system of squirmer microswimmers moving under the influence of gravity at low Reynolds numbers. In addition, the squirmers are…

Fluid Dynamics · Physics 2020-05-13 Felix Rühle , Holger Stark

Microorganisms can preferentially orient and move along gradients of a chemo-attractant (i.e., chemotax) while colonies of many microorganisms can collectively undergo complex dynamics in response to chemo-attractants that they themselves…

Biological Physics · Physics 2013-10-30 Enkeleida Lushi , Raymond E. Goldstein , Michael J. Shelley

We developed a mechanical model of spermatozoal swimming in bulk suspensions. We traced the spatiotemporal elastohydrodynamic interactions and found that spermatozoa engaged in self-organisation: flagellar undulatory motion generated a…

Fluid Dynamics · Physics 2023-02-22 Nanami Taketoshi , Toshihiro Omori , Takuji Ishikawa

Ordered phases emerged in active suspensions of polar swimmers are under long-wavelength hydrodynamic mediated instabilities. In this letter, we show that chemical molecules dissolved in aqueous suspensions, as an unavoidable part of most…

Statistical Mechanics · Physics 2019-06-17 Mehrana R. Nejad , Ali Najafi

We investigate theoretically the collective dynamics of a suspension of low Reynolds number swimmers that are confined to two dimensions by a thin fluid film. Our model swimmer is characterized by internal degrees of freedom which locally…

Biological Physics · Physics 2015-05-19 M. Leoni , T. B. Liverpool

The hydrodynamic interactions of a suspension of self-propelled particles are studied using a direct numerical simulation method which simultaneously solves for the host fluid and the swimming particles. A modified version of the "Smoothed…

Soft Condensed Matter · Physics 2013-01-15 John J. Molina , Yasuya Nakayama , Ryoichi Yamamoto

Microorganisms ofter move in confined, disordered environments, where hydrodynamic couplings can modify their transport behavior. Using extensive finite-element simulations, we investigate the dynamics of microswimmers -- modeled as…

Soft Condensed Matter · Physics 2026-03-24 Mirko Residori , Sebastian Aland , Christina Kurzthaler

Collective states of inanimate particles self-assemble through physical interactions and thermal motion. Despite some phenomenological resemblance, including signatures of criticality, the autonomous dynamics that binds motile agents into…

Soft Condensed Matter · Physics 2023-02-01 Xiangzun Wang , Pin-Chuan Chen , Klaus Kroy , Viktor Holubec , Frank Cichos

Systems of motile microorganisms exhibit a multitude of collective phenomena, including motility-induced phase separation and turbulence. Sensing of the environment and adaptation of movement plays an essential role in the emergent…

Soft Condensed Matter · Physics 2024-06-26 Segun Goh , Elmar Westphal , Roland G. Winkler , Gerhard Gompper

Microswimmers in nature often experience spatial gradients of viscosity. In this work we develop theoretical results for the dynamics of active particles, biological or otherwise, swimming through viscosity gradients. We model the active…

Fluid Dynamics · Physics 2019-10-16 Charu Datt , Gwynn J. Elfring

In this Letter, we study the collective behaviour of a large number of self-propelled microswimmers immersed in a fluid. Using unprecedently large-scale lattice Boltzmann simulations, we reproduce the transition to bacterial turbulence. We…

Soft Condensed Matter · Physics 2017-07-17 Joakim Stenhammar , Cesare Nardini , Rupert W. Nash , Davide Marenduzzo , Alexander Morozov

The "squirmer model" is a classical hydrodynamic model for the motion of interfacially-driven microswimmers, such as self-phoretic colloids or volvocine green algae. To date, most studies using the squirmer model have considered spherical…

Soft Condensed Matter · Physics 2023-10-31 Ruben Poehnl , William E. Uspal

Organisms often swim through density stratified fluids. In this Letter, we investigate the dynamics of small active particles swimming in density gradients and report theoretical evidence of taxis as a result of density stratification…

Soft Condensed Matter · Physics 2023-08-02 Vaseem A. Shaik , Gwynn J. Elfring

Bacteria in biofilms form complex structures and can collectively migrate within mobile aggregates, which is referred to as swarming. This behavior is influenced by a combination of various factors, including morphological characteristics…

Soft Condensed Matter · Physics 2024-01-19 Bohan Wu-Zhang , Dmitry A. Fedosov , Gerhard Gompper

We study active agents embedded in bulk or in confinement explicitly considering hydrodynamics and simulating the swimmers via an implementation inspired by the squirmer model. We develop a Dissipative Particle Dynamics scheme for the…