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We aim to identify the control principles governing the adaptable formation of non-equilibrium structures in actomyosin networks. We build a phenomenological model and predict that biasing the energy dissipated by molecular motors should…

Active networks made of biopolymers and motor proteins are valuable bioinspired systems that have been used in the last decades to study the cytoskeleton and its self-organization under mechanical stimulation. Different techniques are…

Soft Condensed Matter · Physics 2023-05-15 Vahid Nasirimarekani , Olinka Ramìrez-Soto , Stefan Karpitschka , Isabella Guido

Living systems often exhibit internal driving: active, molecular processes drive nonequilibrium phenomena such as metabolism or migration. Active gels constitute a fascinating class of internally driven matter, where molecular motors exert…

Biological Physics · Physics 2015-06-15 Jose Alvarado , Michael Sheinman , Abhinav Sharma , Fred C. MacKintosh , Gijsje H. Koenderink

Developing tissues need to pattern themselves in space and time. A prevalent mechanism to achieve this are pulsatile active stresses generated by the actin cytoskeleton. Active gel theory is a powerful tool to model the dynamics of…

Biological Physics · Physics 2023-02-21 Eloy Merlijn de Kinkelder , Elisabeth Fischer-Friedrich , Sebastian Aland

Complex interactions between cellular systems and their surrounding extracellular matrices are emerging as important mechanical regulators of cell functions such as proliferation, motility, and cell death, and such cellular systems are…

Soft Condensed Matter · Physics 2021-04-16 Emmanuel L. C. VI M. Plan , Julia M. Yeomans , Amin Doostmohammadi

We consider a continuum model of active viscoelastic matter, whereby an active nematic liquid-crystal is coupled to a minimal model of polymer dynamics with a viscoelastic relaxation time $\tau_C$. To explore the resulting interplay between…

Soft Condensed Matter · Physics 2016-03-30 E. J. Hemingway , M. E. Cates , S. M. Fielding

The composite cytoskeleton, comprising interacting networks of semiflexible actin and rigid microtubules, actively generates forces and restructures using motor proteins such as myosins to enable key mechanical processes including cell…

Pattern formation and the mechanics of a mixture of actin filaments and myosin motors that is confined by a rigid membrane is investigated. By using a coarse-grained molecular dynamics model, we demonstrate that the competition between the…

Soft Condensed Matter · Physics 2023-04-12 Mitsusuke Tarama , Tatsuo Shibata

The mechanical properties of cells are dominated by the cytoskeleton, an interconnected network of long elastic filaments. The connections between the filaments are provided by crosslinking proteins, which constitute, next to the filaments,…

Soft Condensed Matter · Physics 2012-10-09 Claus Heussinger

Amorphous elastomers exhibit significant rate-stiffening and unique viscous flow characteristics across a wide range of strain rates, often undergoing glass transition above a strain rate threshold. We have developed a…

Soft Condensed Matter · Physics 2026-05-26 Bibekananda Datta , Sushan Nakarmi , Nitin P. Daphalapurkar

Physical experiments can characterize the elastic response of granular materials in terms of macroscopic state-variables, namely volume (packing) fraction and stress, while the microstructure is not accessible and thus neglected. Here, by…

Soft Condensed Matter · Physics 2015-06-16 Nishant Kumar , Stefan Luding , Vanessa Magnanimo

Mechanically induced folding of passive cross-linkers is a fundamental biological phenomenon. A typical example is a conformational change in myosin II responsible for the power-stroke in skeletal muscles. In this paper we present an…

Biological Physics · Physics 2017-09-14 Matthieu Caruel , Jean-Marc Allain , Lev Truskinovsky

We study a reconstituted composite system consisting of an active microtubule network interdigitated with a passive network of entangled F-actin filaments. Increasing viscoelasticity of the F-actin network controls the emergent dynamics,…

Soft Condensed Matter · Physics 2022-02-16 John Berezney , Bruce L. Goode , Seth Fraden , Zvonimir Dogic

The cytoskeleton relies on diverse populations of motors, filaments, and binding proteins acting in concert to enable non-equilibrium processes ranging from mitosis to chemotaxis. Its versatile reconfigurability, programmed by interactions…

We propose and analyse the properties of a new class of models for the electromechanics of cardiac tissue. The set of governing equations consists of nonlinear elasticity using a viscoelastic and orthotropic exponential constitutive law…

Numerical Analysis · Mathematics 2022-04-08 Adrienne Propp , Alessio Gizzi , Francesc Levrero-Florencio , Ricardo Ruiz-Baier

We present a constitutive model for the passive and active response of skeletal muscles. At variance with more classical approaches, the model is developed exploiting adapted Voigt representations of strain and stress tensors within the…

Soft Condensed Matter · Physics 2026-03-23 Sara Galasso , Giulio G. Giusteri

The actin cytoskeleton is remarkably adaptable and multifunctional. It often organizes into nematic bundles such as contractile rings or stress fibers. However, how a uniform and isotropic actin gel self-organizes into dense nematic bundles…

Inspired by active shape morphing in developing tissues and biomaterials, we investigate two generic mechanochemical models where the deformations of a thin elastic sheet are driven by, and in turn affect, the concentration gradients of a…

Soft Condensed Matter · Physics 2021-07-14 Andrei Zakharov , Kinjal Dasbiswas

We study the high-frequency micro-mechanical response of suspensions composed by cardiac and skeletal muscle myosin by optical trapping interferometry. We observe that in low ionic strength solutions upon the addition of magnesium adenosine…

Soft Condensed Matter · Physics 2024-10-15 Pablo Domínguez-García , Jose R. Pinto , Ana Akrap , Sylvia Jeney

We perform molecular dynamic (MD) simulations of frictional non-thermal particles driven by an externally applied shear stress. After the system jams following a transient flow, we probe its mechanical response in order to clarify whether…

Soft Condensed Matter · Physics 2012-02-09 Raffaele Pastore , Massimo Pica Ciamarra , Antonio Coniglio