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In this article, we consider a swimmer (i.e. a self-deformable body) immersed in a fluid, the flow of which is governed by the stationary Stokes equations. This model is relevant for studying the locomotion of microorganisms or micro robots…

Analysis of PDEs · Mathematics 2012-03-19 Jérôme Lohéac , Alexandre Munnier

The swimming of a deformable planar slab in a viscous incompressible fluid is studied on the basis of the Navier-Stokes equations. A continuum of plane wave displacements, symmetric on both sides of the slab and characterized by a…

Fluid Dynamics · Physics 2016-11-08 B. U. Felderhof

A single flexible filament can be actuated to escape from the scallop theorem and generate net propulsion at low Reynolds number. In this work, we study the dynamics of a simple boundary-driven multi-filament swimmer, a two-arm clamshell…

Fluid Dynamics · Physics 2022-03-07 Shiyuan Hu , Jun Zhang , Michael J. Shelley

Elongate animals and robots use undulatory body waves to locomote through diverse environments. Geometric mechanics provides a framework to model and optimize such systems in highly damped environments, connecting a prescribed shape change…

Robotics · Computer Science 2025-10-21 Jianfeng Lin , Tianyu Wang , Baxi Chong , Matthew Fernandez , Zhaochen Xu , Daniel I. Goldman

Cilia and flagella exhibit regular bending waves that perform mechanical work on the surrounding fluid, to propel cellular swimmers and pump fluids inside organisms. Here, we quantify a force-velocity relationship of the beating flagellum,…

Cell Behavior · Quantitative Biology 2016-12-21 Gary S. Klindt , Christian Ruloff , Christian Wanger , Benjamin M. Friedrich

Changes in calcium concentration along the sperm flagellum regulate sperm motility and hyperactivation, characterized by an increased flagellar bend amplitude and beat asymmetry, enabling the sperm to reach and penetrate the ovum (egg). The…

Numerical Analysis · Mathematics 2019-10-21 Lucia Carichino , Sarah D. Olson

Self-propelled phoretic swimmers are generally studied in the laminar flow regime, where their low speed renders inertial effects negligible and trajectories highly predictable. This research tackles the challenge of propulsion in the…

Fluid Dynamics · Physics 2026-01-19 Alessandro Foradori , Paolo Bettotti

Many swimming microorganisms, such as bacteria and sperm, use flexible flagella to move through viscoelastic media in their natural environments. In this paper we address the effects a viscoelastic fluid has on the motion and beating…

Soft Condensed Matter · Physics 2009-11-13 Henry C. Fu , Charles W. Wolgemuth , Thomas R. Powers

We determine the parametric hull of a given volume which minimizes the total water resistance for a given speed of the ship. The total resistance is the sum of Michell's wave resistance and of the viscous resistance, approximated by…

Optimization and Control · Mathematics 2014-10-13 Julien Dambrine , Morgan Pierre , Germain Rousseaux

The intricate wobbling motion of flagellated bacteria, characterized by the periodic precession of the cell body, is a determinant factor in their motility and navigation within complex fluid environments. While well-studied in quiescent…

Soft Condensed Matter · Physics 2026-01-26 Wei Feng , Fanglong Dang , Hao Luo , Alan C. H. Tsang , Yanan Liu , Guangyin Jing

Active swimmers are ubiquitous in nature, found in many diverse biological systems ranging from bacteria to vertebrate fish. Of particular importance are sperm cells which are swimmers that are crucial for the survival of many species…

Biological Physics · Physics 2017-11-15 James L. Kingsley , Utkan Demirci , Erkan Tuzel

Fish swim with flexible fins that stand in stark contrast to the rigid propulsors of engineered vehicles. Using numerical simulations of the dynamics of flow-structure interaction, we have found that dorso-ventral deformation in flexible…

Fluid Dynamics · Physics 2026-04-16 Sushrut Kumar , Matthew J. McHenry , Jung-Hee Seo , Rajat Mittal

The hydrodynamic interactions among bacterial cell bodies, flagella, and surrounding boundaries are essential for understanding bacterial motility in complex environments. In this study, we demonstrate that each slender flagellum can be…

Soft Condensed Matter · Physics 2025-01-07 Baopi Liu , Lu Chen , Ji Zhang

We derive a theorem for the lower bound on the energy dissipation rate by a rigid surface-driven active microswimmer of arbitrary shape in a fluid at low Reynolds number. We show that, for any swimmer, the minimum dissipation at a given…

Fluid Dynamics · Physics 2021-01-27 Babak Nasouri , Andrej Vilfan , Ramin Golestanian

Many microorganisms propel through complex media by deformations of their flagella. The beat is thought to emerge from interactions between forces of the surrounding fluid, passive elastic response from deformations of the flagellum, and…

Fluid Dynamics · Physics 2024-01-23 Kathryn G. Link , Robert D. Guy , Becca Thomases , Paulo E. Arratia

Swimming cells and microorganisms are as diverse in their collective dynamics as they are in their individual shapes and propulsion mechanisms. Even for sperm cells, which have a stereotyped shape consisting of a cell body connected to a…

Fluid Dynamics · Physics 2018-06-25 S. F. Schoeller , E. E. Keaveny

In this paper we study a mathematical model of one-dimensional swimmers performing a planar motion while fully immersed in a viscous fluid. The swimmers are assumed to be of small size, and all inertial effects are neglected. Hydrodynamic…

Optimization and Control · Mathematics 2014-04-25 Gianni Dal Maso , Antonio DeSimone , Marco Morandotti

In a recent letter (Friedrich et al., Phys. Rev. Lett. 109:138102, 2012), a minimal model swimmer was proposed that propels itself at low Reynolds numbers by a revolving motion of a pair of spheres. The motion of the two spheres can…

Cell Behavior · Quantitative Biology 2015-06-12 Katja Polotzek , Benjamin M. Friedrich

We present a unified discussion of three types of near-spherical amoeboid microswimmers, driven by periodic, axially symmetric, achiral deformations (swim strokes): a solid deformable body, a vesicle with incompressible fluid membrane, and…

Soft Condensed Matter · Physics 2025-02-13 Reiner Kree , Annette Zippelius

We investigate the optimal shapes of the hydrodynamic resistance of a rigid body set in motion in a Stokes flow. In this low Reynolds number regime, the hydrodynamic drag properties of an object are encoded in a finite number of parameters…

Fluid Dynamics · Physics 2022-07-14 Clment Moreau , Kenta Ishimoto , Yannick Privat
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