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Hydrodynamic synchronization provides a general mechanism for the spontaneous emergence of coherent beating states in independently driven mesoscopic oscillators. A complete physical picture of those phenomena is of definite importance to…

Soft Condensed Matter · Physics 2014-01-29 Nick Koumakis , Roberto Di Leonardo

We explore in this paper the phenomenon of photofocusing: a coupling between flow vorticity and biased swimming of microalgae toward a light source that produces a focusing of the microswimmer suspension. We combine experiments that…

Soft Condensed Matter · Physics 2016-05-25 Matthieu Martin , Alexandre Barzyk , Eric Bertin , Philippe Peyla , Salima Rafai

Three-dimensional simulations with fully resolved hydrodynamics are performed to study the collective motion of model swimmers in confinement. We show that certain swimming mechanisms can lead to traveling wave-like collective motion even…

Soft Condensed Matter · Physics 2016-04-20 Norihiro Oyama , John Jairo Molina , Ryoichi Yamamoto

We study the fluid dynamics of two fish-like bodies with synchronised swimming patterns. Our studies are based on two-dimensional simulations of viscous incompressible flows. We distinguish between motion patterns that are externally…

In biology, cells undergo deformations under the action of flow caused by the fluid surrounding them. These flows lead to shape changes and instabilities that have been explored in detail for single component vesicles. However, cell…

Fluid Dynamics · Physics 2025-10-15 Anirudh Venkatesh , Vivek Narsimhan

Sperm are propelled by bending waves travelling along the flagellum. During steering in gradients of sensory cues, sperm adjust the flagellar beat waveform. Symmetric and asymmetric beat waveforms produce straight and curved swimming paths,…

Within the RSJ model, we performed a theoretical analysis of phase-locking in elementary strongly coupled Josephson junction cells. For this purpose, we developed a systematic method allowing the investigation of phase-locking in cells with…

Condensed Matter · Physics 2016-08-31 M. Basler , W. Krech , K. Yu. Platov

Cilia and flagella in biological systems often show large scale cooperative behaviors such as the synchronization of their beats in "metachronal waves". These are beautiful examples of emergent dynamics in biology, and are essential for…

Soft Condensed Matter · Physics 2016-04-20 Nicolas Bruot , Pietro Cicuta

Suspensions of swimming micro-organisms are known to undergo intricate collective dynamics as a result of hydrodynamic and collision interactions. Micro-swimmers, such as bacteria and micro-algae, naturally live and have evolved in complex…

Fluid Dynamics · Physics 2025-10-31 Yasser Almoteri , Enkeleida Lushi

Swimming cells and microorganisms must often move though complex fluids that contain an immersed microstructure such as polymer molecules, or filaments. In many important biological processes, such as mammalian reproduction and bacterial…

Fluid Dynamics · Physics 2018-08-06 Arshad Kamal , Eric E Keaveny

Cilia and flagella exhibit regular bending waves that perform mechanical work on the surrounding fluid, to propel cellular swimmers and pump fluids inside organisms. Here, we quantify a force-velocity relationship of the beating flagellum,…

Cell Behavior · Quantitative Biology 2016-12-21 Gary S. Klindt , Christian Ruloff , Christian Wanger , Benjamin M. Friedrich

We show that spontaneous density segregation in dense systems of aligning circle swimmers is a condensation phenomenon at odds with the phase separation scenarios usually observed in two-dimensional active matter. The condensates, which…

Soft Condensed Matter · Physics 2024-12-20 Yujia Wang , Bruno Ventéjou , Hugues Chaté , Xia-qing Shi

Microswimmers are encountered in a wide variety of biophysical settings. When interacting with flow fields, they show interesting dynamical features such as trapping, clustering, and preferential orientation. One important step towards the…

Fluid Dynamics · Physics 2020-10-02 José-Agustín Arguedas-Leiva , Michael Wilczek

Cilia and flagella are actively bending slender organelles, performing functions such as motility, feeding and embryonic symmetry breaking. We review the mechanics of viscous-dominated microscale flow, including time-reversal symmetry, drag…

Quantitative Methods · Quantitative Biology 2013-09-06 Thomas D. Montenegro-Johnson , Andrew A. Smith , David J. Smith , Daniel Loghin , John R. Blake

We study the effect of a nearby planar wall on the propulsion of a spherical phoretic micro-swimmer driven by reactions on its surface. An asymmetric coverage of catalysts on its surface which absorb reactants and generate products gives…

Soft Condensed Matter · Physics 2017-06-08 Yahaya Ibrahim , Tanniemola B. Liverpool

We analyze a minimal model for a rigid spherical microswimmer and explore the consequences of its extended surface on the interplay between its self-propulsion and flow properties. The model is the first order representation of…

Soft Condensed Matter · Physics 2017-12-06 Tapan Chandra Adhyapak , Sara Jabbari-Farouji

Self-organized dynamic patterns in dense active matter are striking manifestations of non-equilibrium physics. A prominent example is the macroscopic elliptical motion observed in quasi-2D bacterial suspensions, which has lacked a physical…

Soft Condensed Matter · Physics 2026-03-30 Bing Miao , Lei-Han Tang

We combine a general formulation of microswimmmer equations of motion with a numerical bead-shell model to calculate the hydrodynamic interactions with the fluid, from which the swimming speed, power and efficiency are extracted. From this…

Soft Condensed Matter · Physics 2017-03-07 Bram Bet , Gijs Boosten , Marjolein Dijkstra , René van Roij

Efficient locomotion is important for the evolution of complex life, yet the physical principles selecting specific swimming strokes often remain entangled with biological constraints. In viscous fluids, the scallop theorem constrains the…

Biological Physics · Physics 2026-03-11 Takahiro Kanazawa , Kenta Ishimoto , Kyogo Kawaguchi

In active systems, whose constituents have non-equilibrium dynamics at local level, fluid-fluid phase separation is widely observed. Examples include the formation of membraneless organelles within cells; the clustering of self-propelled…

Soft Condensed Matter · Physics 2025-07-11 M. E. Cates , C. Nardini
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