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The vast majority of mutations are deleterious, and are eliminated by purifying selection. Yet in finite asexual populations, purifying selection cannot completely prevent the accumulation of deleterious mutations due to Muller's ratchet:…

Populations and Evolution · Quantitative Biology 2011-10-14 Sidhartha Goyal , Daniel J. Balick , Elizabeth R. Jerison , Richard A. Neher , Boris I. Shraiman , Michael M. Desai

We investigate the exploration of rugged fitness landscapes by spatially structured populations with demes on the nodes of a graph, connected by migrations. In the rare migration regime, we find that finite structures can adapt more…

Populations and Evolution · Quantitative Biology 2025-02-07 Richard Servajean , Arthur Alexandre , Anne-Florence Bitbol

The quantitative characterization of mutational landscapes is a task of outstanding importance in evolutionary and medical biology: It is, e.g., of central importance for our understanding of the phenotypic effect of mutations related to…

Quantitative Methods · Quantitative Biology 2015-10-13 Matteo Figliuzzi , Hervé Jacquier , Alexander Schug , Olivier Tenaillon , Martin Weigt

We revisit the model by Wiser, Ribeck, and Lenski (Science \textbf{342} (2013), 1364--1367), which describes how the mean fitness increases over time due to beneficial mutations in Lenski's long-term evolution experiment. We develop the…

Populations and Evolution · Quantitative Biology 2020-02-21 Ellen Baake , Adrián González Casanova , Sebastian Probst , Anton Wakolbinger

Muller's ratchet describes the irreversible accumulation of deleterious mutations in asexual populations. In well-mixed populations the speed of fitness decline is exponentially small in the population size, and any positive rate of…

Populations and Evolution · Quantitative Biology 2018-09-26 Su-Chan Park , Philipp Klatt , Joachim Krug

With a view to connecting random mutation on the molecular level to punctuated equilibrium behavior on the phenotype level, we propose a new model for biological evolution, which incorporates random mutation and natural selection. In this…

Condensed Matter · Physics 2009-10-28 M. Y. Choi , H. Y. Lee , D. Kim , S. H. Park

Phenotype of biological systems needs to be robust against mutation in order to sustain themselves between generations. On the other hand, phenotype of an individual also needs to be robust against fluctuations of both internal and external…

Populations and Evolution · Quantitative Biology 2015-06-26 Kunihiko Kaneko

Epistatic interactions between residues determine a protein's adaptability and shape its evolutionary trajectory. When a protein experiences a changed environment, it is under strong selection to find a peak in the new fitness landscape. It…

Populations and Evolution · Quantitative Biology 2016-04-01 Aditi Gupta , Christoph Adami

Fitness landscapes provide a quantitative framework for understanding how natural selection shapes evolutionary trajectories. A central feature of these landscapes is their number of local optima, which determines whether fitness-increasing…

Populations and Evolution · Quantitative Biology 2026-05-06 Mahan Ghafari , Alejandro Castro Cabrera , Alejandro Lage-Castellanos , Guillaume Achaz , Joachim Krug , Luca Ferretti

When polygenic traits are under stabilizing selection, many different combinations of alleles allow close adaptation to the optimum. If alleles have equal effects, all combinations that result in the same deviation from the optimum are…

Populations and Evolution · Quantitative Biology 2014-04-04 Harold P. de Vladar , Nick Barton

Phenotypic evolution implies sequential fixations of new genomic sequences. The speed at which these mutations fixate depends, in part, on the relative fitness (selection coefficient) of the mutant vs. the ancestor. Using a simple…

Populations and Evolution · Quantitative Biology 2014-02-04 Sorin Tanase-Nicola , Ilya Nemenman

We study the evolution of asexual microorganisms with small mutation rate in fluctuating environments, and develop techniques that allow us to expand the formal solution of the evolution equations to first order in the mutation rate. Our…

Biological Physics · Physics 2009-11-06 Claus Wilke , Christopher Ronnewinkel

Proteins evolve through complex sequence spaces, with fitness landscapes serving as a conceptual framework that links sequence to function. Fitness landscapes can be smooth, where multiple similarly accessible evolutionary paths are…

Populations and Evolution · Quantitative Biology 2024-11-21 Mahakaran Sandhu , John Chen , Dana Matthews , Matthew A Spence , Sacha B Pulsford , Barnabas Gall , James Nichols , Nobuhiko Tokuriki , Colin J Jackson

When mutations are rampant, quasispecies theory or Eigen's model predicts that the fittest type in a population may not dominate. Beyond a critical mutation rate, the population may even be delocalized completely from the peak of the…

Populations and Evolution · Quantitative Biology 2026-04-15 C. J. Palpal-latoc , Ian Vega

We considered a {multi-block} molecular model of biological evolution, in which fitness is a function of the mean types of alleles located at different parts (blocks) of the genome. We formulated an infinite population model with selection…

Populations and Evolution · Quantitative Biology 2015-06-12 David B. Saakian , Zara Kirakosyan , Chin-Kun Hu

Due to stochastic fluctuations arising from finite population size, known as genetic drift, the ability of a population to explore a rugged fitness landscape depends on its size. In the weak mutation regime, while the mean steady-state…

Populations and Evolution · Quantitative Biology 2024-12-30 Richard Servajean , Anne-Florence Bitbol

We propose a model to characterize how a diffusing population adapts under a time periodic selection, while its environment undergoes shifts and size changes, leading to significant differences with classical results on fixed domains. After…

Analysis of PDEs · Mathematics 2025-06-05 Matthieu Alfaro , Adel Blouza , Nessim Dhaouadi

The adaptive evolution of large asexual populations is generally characterized by competition between clones carrying different beneficial mutations. This interference phenomenon slows down the adaptation speed and makes the theoretical…

Populations and Evolution · Quantitative Biology 2012-12-20 Maria Rita Fumagalli , Matteo Osella , Philippe Thomen , Francois Heslot , Marco Cosentino Lagomarsino

We investigate the evolutionary dynamics of a finite population of RNA sequences adapting to a neutral fitness landscape. Despite the lack of differential fitness between viable sequences, we observe typical properties of adaptive…

Populations and Evolution · Quantitative Biology 2007-05-23 Robert Forster , Christoph Adami , Claus O. Wilke

Phenotypically structured equations arise in population biology to describe the interaction of species with their environment that brings the nutrients. This interaction usually leads to selection of the fittest individuals. Models used in…

Analysis of PDEs · Mathematics 2015-06-18 Alexander Lorz , Benoit Perthame
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