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We study a class of the stochastic May-Leonard models, with three species dominating each other in a cyclic nonhierarchical way, according to the rock-paper-scissors game. We introduce an unevenness in the system, by considering that one of…

Populations and Evolution · Quantitative Biology 2019-05-23 J. Menezes , B. Moura , T. A. Pereira

The Jungle Game is used in population dynamics to describe cyclic competition among species that interact via a food chain. The dynamics of the Jungle Game supports a heteroclinic network whose cycles represent coexisting species. The…

Dynamical Systems · Mathematics 2024-01-17 Sofia B. S. D. Castro , Ana M. J. Ferreira , Isabel S. Labouriau

We study a three-species cyclic game system where organisms face a contagious disease whose virulence may change by a pathogen mutation. As a responsive defence strategy, organisms' mobility is restricted to reduce disease dissemination in…

Populations and Evolution · Quantitative Biology 2022-12-27 J. Menezes , S. Batista , E. Rangel

The spatial segregation of species is fundamental to ecosystem formation and stability. Behavioural strategies may determine where species are located and how their interactions change the local environment arrangement. In response to…

Populations and Evolution · Quantitative Biology 2021-03-23 B. Moura , J. Menezes

We study the cyclic dominance of three species in two-dimensional constrained Newman-Watts networks with a four-state variant of the rock-paper-scissors game. By limiting the maximal connection distance $R_{max}$ in Newman-Watts networks…

Populations and Evolution · Quantitative Biology 2009-07-03 Guo-Yong Zhang , Yong Chen , Wei-Kai Qi , Shao-Meng Qin

In a wide variety of natural systems, closely-related microbial strains coexist stably, resulting in high levels of fine-scale biodiversity. However, the mechanisms that stabilize this coexistence are not fully understood. Spatial…

Populations and Evolution · Quantitative Biology 2023-03-29 Casey Barkan , Shenshen Wang

Antipredator behavior is present in many biological systems where individuals collectively react to an imminent attack. The antipredator response may influence spatial pattern formation and ecosystem stability but requires an organism's…

Populations and Evolution · Quantitative Biology 2021-11-10 J. Menezes , B. Moura

The rock-paper-scissor game -- which is characterized by three strategies R,P,S, satisfying the non-transitive relations S excludes P, P excludes R, and R excludes S -- serves as a simple prototype for studying more complex non-transitive…

Populations and Evolution · Quantitative Biology 2015-12-16 Sebastian J. Schreiber , Timothy P. Killingback

Computer simulations of minimal population-dynamics models have long been used to explore questions in ecosystems coexistence and species biodiversity, via simple agent-based models of three interacting species, referred to as $R$, $P$, and…

Populations and Evolution · Quantitative Biology 2024-10-07 Dave Cliff

The classical game of rock-paper-scissors have inspired experiments and spatial model systems that address robustness of biological diversity. In particular the game nicely illustrates that cyclic interactions allow multiple strategies to…

Cellular Automata and Lattice Gases · Physics 2016-04-20 Namiko Mitarai , Ivar Gunnarson , Buster Niels Pedersen , Christian Anker Rosiek , Kim Sneppen

How diversity is maintained in natural ecosystems is a long-standing question in Theoretical Ecology. By studying a system that combines ecological dynamics, heterogeneous interactions and spatial structure, we uncover a new mechanism for…

Populations and Evolution · Quantitative Biology 2024-03-21 Giulia Garcia Lorenzana , Ada Altieri , Giulio Biroli

Disease outbreaks affect many ecosystems threatening species that also fight against other natural enemies. We investigate a cyclic game system with $5$ species, whose organisms outcompete according to the rules of a generalised spatial…

Populations and Evolution · Quantitative Biology 2022-05-13 E. Rangel , B. Moura , J. Menezes

Natural birth and death are fundamental mechanisms of population dynamics in ecosystems and have played pivotal roles in shaping population dynamics. Nevertheless, in studies of cyclic competition systems governed by the rock-paper-scissors…

Populations and Evolution · Quantitative Biology 2026-02-19 Yikang Lu , Wenhao She , Xiaofang Duan , Junpyo Park

Species extinction occurs regularly and unavoidably in ecological systems. The time scales for extinction can broadly vary and inform on the ecosystem's stability. We study the spatio-temporal extinction dynamics of a paradigmatic…

Populations and Evolution · Quantitative Biology 2011-03-02 S. Rulands , T. Reichenbach , E. Frey

When three species compete cyclically in a well-mixed, stochastic system of $N$ individuals, extinction is known to typically occur at times scaling as the system size $N$. This happens, for example, in rock-paper-scissors games or…

Physics and Society · Physics 2019-02-20 Kevin E. Bassler , Erwin Frey , R. K. P. Zia

We study the generalised rock-paper-scissors game with five species whose organisms face local epidemic outbreaks. As an evolutionary behavioural survival strategy, organisms of one out of the species move in the direction with more enemies…

Populations and Evolution · Quantitative Biology 2022-12-29 J. Menezes , B. Moura , E. Rangel

Species survival in the $(3, 1)$ May-Leonard system is determined by the mobility, with a critical mobility threshold between long-term coexistence and extinction. We show experimentally that the critical mobility threshold is determined by…

Populations and Evolution · Quantitative Biology 2021-11-09 Mark Lowell

Recurrence plots and their associated quantifiers provide a robust framework for detecting and characterising complex patterns in non-linear time-series. In this paper, we employ recurrence quantification analysis to investigate the…

Populations and Evolution · Quantitative Biology 2025-07-29 M. S. Palmero , M. Bongestab , N. Marwan

The spatial rock-paper-scissors ecosystem, where three species interact cyclically, is a model example of how spatial structure can maintain biodiversity. We here consider such a system for a broad range of interaction rates. When one…

Biological Physics · Physics 2013-06-21 Jeppe Juul , Kim Sneppen , Joachim Mathiesen

The maintenance of diversity, the `commonness of rarity', and compositional turnover are ubiquitous features of species-rich communities. Through a minimal model, we consider how these features reflect the interplay between environmental…

Populations and Evolution · Quantitative Biology 2025-12-04 Emil Mallmin , Arne Traulsen , Silvia De Monte