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We use a continuum, two-fluid approach to study a mixture of two active nematic fluids. Even in the absence of thermodynamically-driven ordering, for mixtures of different activities we observe turbulent microphase separation, where domains…

Soft Condensed Matter · Physics 2025-02-03 Saraswat Bhattacharyya , Julia M. Yeomans

In tissues as diverse as amphibian skin and the human airway, the cilia that propel fluid are grouped in sparsely distributed multiciliated cells (MCCs). We investigate fluid transport in this "mosaic" architecture, with emphasis on the…

Soft Condensed Matter · Physics 2021-11-17 Francesco Boselli , Jerome Jullien , Eric Lauga , Raymond E. Goldstein

The present work reports the formation and the characterization of antipleptic and symplectic metachronal waves in 3D cilia arrays immersed in a two-fluid environment, with a viscosity ratio of 20. A coupled…

Biological Physics · Physics 2017-08-02 Sylvain Chateau , Julien Favier , Umberto D 'Ortona , Sébastien Poncet

The functions of many networked systems in physics, biology or engineering rely on a coordinated or synchronized dynamics of its constituents. In power grids for example, all generators must synchronize and run at the same frequency and…

Adaptation and Self-Organizing Systems · Physics 2017-09-13 Debsankha Manik , Marc Timme , Dirk Witthaut

Biological systems are robust to perturbations at both the genetic and environmental levels. Yet, these same perturbations can elicit variation in behavior. The interplay between functional robustness and behavioral variability is…

Biological Physics · Physics 2021-06-03 Veikko Geyer , Jonathon Howard , Pablo Sartori

Flows over two side-by-side circular cylinders exhibit fascinating flow physics due to complex interactions between the coupled wakes. However, their mutual interference effects have not been elucidated in a quantitative manner thus far. In…

Fluid Dynamics · Physics 2022-01-05 Kai Zhang , Md. Naimul Haque

Near a solid boundary, E. coli swims in clockwise circular motion. We provide a hydrodynamic model for this behavior. We show that circular trajectories are natural consequences of force-free and torque-free swimming, and the hydrodynamic…

Soft Condensed Matter · Physics 2016-08-31 Eric Lauga , Willow R. DiLuzio , George M. Whitesides , Howard A. Stone

Many aquatic organisms are able to track ambient flow disturbances and locate their source. These tasks are particularly challenging because they require the organism to sense local flow information and respond accordingly. Details of how…

Optimization and Control · Mathematics 2018-04-10 Brendan Colvert , Eva Kanso

We introduce a phenomenological theory for a new class of soft active fluids, with the ability to synchronise. Our theoretical framework describes the macroscopic behaviour of a collection of interacting anisotropic elements with cyclic…

Soft Condensed Matter · Physics 2015-06-16 M. Leoni , T. B. Liverpool

Ciliary flows are generated by a vast array of eukaryotic organisms, from unicellular algae to mammals, and occur in a range of different geometrical configurations. We employ a point torque -- or `rotlet' -- model to capture the…

We present an experimental study of a confined nanoflow, which is generated by a sphere oscillating in the proximity of a flat solid wall in a simple fluid. Varying the oscillation frequency, the confining length scale and the fluid mean…

Fluid Dynamics · Physics 2015-06-03 C. Lissandrello , V. Yakhot , K. L. Ekinci

An active chiral rotor is a spherical object that can generate chiral flows in a fluid by rotating about an axis. For example, if the flow around the upper hemisphere of the chiral rotor is in a clockwise direction, then the flow in the…

Soft Condensed Matter · Physics 2025-01-29 R. Maity , Snigdha Thakur , P. S. Burada

Filaments on the surface of a microorganism such as Paramecium or Ophalina beat highly synchronized and form so-called metachronal waves that travel along the surfaces. In order to study under what principal conditions these waves form, we…

Biological Physics · Physics 2010-12-16 Christopher Wollin , Holger Stark

We study theoretically the behavior of a class of hydrodynamic dipoles. This study is motivated by recent experiments on synthetic and biological swimmers in microfluidic \textit{Hele-Shaw} type geometries. Under such confinement, a…

Fluid Dynamics · Physics 2015-09-30 Eva Kanso , Alan Tsang

We present the hydrodynamic theory of active XY spins coupled with flow fields, for systems both having and or lacking number conservation in two dimensions (2D). For the latter, with strong activity or nonequilibrium drive, the system can…

Statistical Mechanics · Physics 2019-11-04 Astik Haldar , Abhik Basu

The flow field generated by a swimming bacterium serves as a fundamental building block for understanding hydrodynamic interactions between bacteria. Although the flow field generated by a force dipole (stresslet) well captures the fluid…

Soft Condensed Matter · Physics 2026-03-27 Yaochen Yang , Daiki Matsunaga , Da Wei , Fanlong Meng

We present a theory for self-driven fluids, such as motorized cytoskeletal extracts or bacterial suspensions, that takes into account the underlying periodic duty cycle carried by the active particles of which the system is composed. We…

Soft Condensed Matter · Physics 2013-12-09 Sebastian Fürthauer , Sriram Ramaswamy

Cilia are ubiquitous organelles involves in eukaryotic motility. They are long, slender, and motile protrusions from the cell body. They undergo active regular oscillatory beating patterns that can propel cells, such as the algae…

Subcellular Processes · Quantitative Biology 2019-05-13 Pablo Sartori

Despite evidence for a hydrodynamic origin of flagellar synchronization between different eukaryotic cells, recent experiments have shown that in single multi-flagellated organisms, coordination hinges instead on direct basal body…

Biological Physics · Physics 2018-10-12 Yujie Liu , Rory Claydon , Marco Polin , Douglas R. Brumley

We investigate the hydrodynamic interaction between two elastic swimmers which are composed of three spheres and two harmonic springs. In this model, the natural length of each spring is assumed to undergo a prescribed cyclic change,…

Soft Condensed Matter · Physics 2019-05-22 Mizuki Kuroda , Kento Yasuda , Shigeyuki Komura
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