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Liquid-liquid phase separation is important across biology, physics, and materials science. Although usually studied at equilibrium, active components - such as motor proteins, enzymes, and synthetic microswimmers - are increasingly…

Soft Condensed Matter · Physics 2026-01-06 Kan Chang , Yulin Li , Ming Yuan , Masaki Sano , Zhihong You , H. P. Zhang

The motility of cilia and flagella is driven by thousands of dynein motors that hydrolyze adenosine triphosphate (ATP). Despite decades of genetic, biochemical, structural and biophysical studies, some aspects of ciliary motility remain…

Biological Physics · Physics 2015-12-22 Daniel T. N. Chen , Michael Heymann , Seth Fraden , Daniela Nicastro , Zvonimir Dogic

Elastic confinements are an important component of many biological systems and dictate the transport properties of suspended particles under flow. In this chapter, we review the Brownian motion of a particle moving in the vicinity of a…

Soft Condensed Matter · Physics 2022-10-28 Abdallah Daddi-Moussa-Ider , Stephan Gekle

Biological systems are robust to perturbations at both the genetic and environmental levels. Yet, these same perturbations can elicit variation in behavior. The interplay between functional robustness and behavioral variability is…

Biological Physics · Physics 2021-06-03 Veikko Geyer , Jonathon Howard , Pablo Sartori

The bacterial flagellar motor drives the rotation of flagellar filaments and enables many species of bacteria to swim. Torque is generated by interaction of stator units, anchored to the peptidoglycan cell wall, with the rotor. Recent…

Subcellular Processes · Quantitative Biology 2010-08-18 G. Meacci , Y. Tu

The acoustofluidic method holds great promise for manipulating microorganisms. When exposed to the steady vortex structures of acoustic streaming flow, these microorganisms exhibit intriguing dynamic behaviors, such as hydrodynamic trapping…

Fluid Dynamics · Physics 2025-04-25 Xuyang Sun , Wenchang Tan , Yi Man

Motivated by the motion of biopolymers and membranes in solution, this article presents a formulation of the equations of motion for curves and surfaces in a viscous fluid. We focus on geometrical aspects and simple variational methods for…

Soft Condensed Matter · Physics 2010-05-26 Thomas R. Powers

Some microorganisms, such as spermatozoa, synchronize their flagella when swimming in close proximity. Using a simplified model (two infinite, parallel, two-dimensional waving sheets), we show that phase-locking arises from hydrodynamics…

Biological Physics · Physics 2009-09-15 Gwynn J. Elfring , Eric Lauga

We present a simplified theory for semiflexible flagella under the action of a traveling-wave perturbation that emulates the organized active forces generated by molecular motors, capable of inducing beating patterns to the filament. By…

Soft Condensed Matter · Physics 2025-07-16 Fabio Cecconi , Dario Lucente , Andrea Puglisi , Massimiliano Viale

Exploiting the "natural" frame of space curves, we formulate an intrinsic dynamics of twisted elastic filaments in viscous fluids. A pair of coupled nonlinear equations describing the temporal evolution of the filament's complex curvature…

Soft Condensed Matter · Physics 2009-10-31 Raymond E. Goldstein , Thomas R. Powers , Chris H. Wiggins

Microorganisms such as algae and bacteria move in a viscous environment with extremely low Reynolds ($Re$), where the viscous drag dominates the inertial forces. They have adapted to this environment by developing specialized features such…

Robotics · Computer Science 2024-12-10 Nnamdi Chikere , Yasemin Ozkan-Aydin

We study the fully-coupled dynamics between a fully-developed turbulent flow and an ensemble of immersed flexible fibers. We vary the concentration of the suspension, the mechanical properties and the length of the fibers in a vast…

Fluid Dynamics · Physics 2021-08-24 Stefano Olivieri , Andrea Mazzino , Marco E. Rosti

The stochastic dynamics of flagellar beating for micro-swimmers, such as flagellated cells, sperms and microalgae, is widely thought to include a feedback mechanism between flagellar shape and the rate of activation/de-activation of the $N…

Soft Condensed Matter · Physics 2026-05-01 F. Fanelli , A. Puglisi

The slime mould Physarum polycephalum has emerged as a model for self-organisation and coordination of contractile activity at large spatial scales. This self-organisation largely results from cytoplasmic flows generated by propagating…

Soft Condensed Matter · Physics 2025-09-23 Raphael Saiseau , Valentin Busson , Marc Durand

We analyse the motion of a flagellated bacterium in a two-fluid medium using slender body theory. The two-fluid model is useful for describing a body moving through a complex fluid with a microstructure whose length scale is comparable to…

Fluid Dynamics · Physics 2024-12-11 Sabarish V. Narayanan , Donald L. Koch , Sarah Hormozi

To survive in harsh conditions, motile bacteria swim in complex environment and respond to the surrounding flow. Here we develop a PDE model describing how the flagella bending affects macroscopic properties of bacterial suspensions. First,…

Analysis of PDEs · Mathematics 2016-10-11 Mykhailo Potomkin , Magali Tournus , Leonid Berlyand , Igor Aranson

Flagellar-driven locomotion plays a critical role in bacterial attachment and colonization of surfaces, contributing to the risks of contamination and infection. Tremendous attempts to uncover the underlying principles governing bacterial…

Soft Condensed Matter · Physics 2025-02-19 Xin-Xin Xu , Yangguang Tian , Yuhe Pu , Bingchen Che , Hao Luo , Yanan Liu , Yan-Jun Liu , Guangyin Jing

Thiovulum majus, which is one of the fastest known bacteria, swims using hundreds of flagella. Unlike typical pusher cells, which swim in circular paths over hard surfaces, a T. majus cell turns its flagella normal to the surface. To probe…

Biological Physics · Physics 2023-06-21 Alexander P Petroff , Schuyler McDonough

The accumulation of swimming bacteria near surfaces may lead to biological processes such as biofilm formation and wound infection. Previous experimental observations of Vibrio alginolyticus showed an interesting correlation between the…

Biological Physics · Physics 2023-07-04 Vahid Nourian , Henry Shum

Efficient bacterial chromosome segregation typically requires the coordinated action of a three-components, fueled by adenosine triphosphate machinery called the partition complex. We present a phenomenological model accounting for the…