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Biological cells sense and respond to mechanical forces, but how such a mechanosensing proccess takes place in a nonlinear inhomogeneous fibrous matrix remains unknown. We show that cells in a fibrous matrix induce deformation fields that…

Biological Physics · Physics 2015-06-10 Jacob Notbohm , Ayelet Lesman , Phoebus Rosakis , David A. Tirrell , Guruswami Ravichandran

A hallmark of biological tissues, viewed as complex cellular materials, is the active generation of mechanical stresses by cellular processes, such as cell divisions. Each cellular event generates a force dipole that deforms the surrounding…

The dynamics of cellular pattern formation is crucial for understanding embryonic development and tissue morphogenesis. Recent studies have shown that human dermal fibroblasts cultured on liquid crystal elastomers can exhibit an increase in…

Biological Physics · Physics 2023-08-25 Mengyang Gu , Xinyi Fang , Yimin Luo

Cells and other soft particles are often forced to flow in confined geometries in both laboratory and natural environments, where the elastic deformation induces an additional drag and pressure drop across the particle. In contrast with…

Fluid Dynamics · Physics 2025-07-09 Charles Paul Moore , Hiba Belkadi , Brouna Safi , Gabriel Amselem , Charles N. Baroud

In biological settings membranes typically interact locally with other membranes or the extracellular matrix in the exterior, as well as with internal cellular structures such as the cytoskeleton. Characterization of the dynamic properties…

Biological Physics · Physics 2019-09-04 Josip Augustin Janeš , Daniel Schmidt , Udo Seifert , Ana-Sunčana Smith

The collective action of actively contractile units embedded in elastic biopolymer networks plays a crucial role in regulating the network's macroscopic mechanical response. Here, we investigate how the macroscopic boundary stress in model…

Soft Condensed Matter · Physics 2026-03-25 Abhinav Kumar , David A. Quint , Kinjal Dasbiswas

Optimal characterization of the mechanical properties of both cells and their surrounding is an issue of major interest. Indeed, cell function and development are strongly influenced by external stimuli. Furthermore, a change in cell…

We investigate the mechanical interplay between the spatial organization of the actin cytoskeleton and the shape of animal cells adhering on micropillar arrays. Using a combination of analytical work, computer simulations and in vitro…

Within developing embryos, tissues flow and reorganize dramatically on timescales as short as minutes. This includes epithelial tissues, which often narrow and elongate in convergent extension movements due to anisotropies in external…

Tissues and Organs · Quantitative Biology 2020-06-24 Xun Wang , Matthias Merkel , Leo B. Sutter , Gonca Erdemci-Tandogan , M. Lisa Manning , Karen E. Kasza

Many morphogenetic processes involve mechanical rearrangement of epithelial tissues that is driven by precisely regulated cytoskeletal forces and cell adhesion. The mechanical state of the cell and intercellular adhesion are not only the…

Cell Behavior · Quantitative Biology 2015-06-03 Kevin K. Chiou , Lars Hufnagel , Boris I. Shraiman

Tuning of active prestress e.g. through activity of molecular motors constitutes a powerful cellular tool to adjust cellular stiffness through nonlinear material properties. Understanding this tool is an important prerequisite for our…

Soft Condensed Matter · Physics 2018-08-24 Elisabeth Fischer-Friedrich

Actin and myosin drive many instances of force generation, deformation, and shape change in cells, tissues, and organisms. In particular, cytoskeletal actomyosin is remarkable in its adaptive architecture, responding to a host of…

Soft Condensed Matter · Physics 2025-12-18 James Clarke , Hyunjae Lee , Kyla Wong , Julia Glenn , Aniket Marne , Yoichi Miyahara , José Alvarado

Cell layers eliminate unwanted cells through the extrusion process, which underlines healthy versus flawed tissue behaviors. Although several biochemical pathways have been identified, the underlying mechanical basis including the forces…

Soft Condensed Matter · Physics 2023-04-20 Siavash Monfared , Guruswami Ravichandran , Jose E. Andrade , Amin Doostmohammadi

Active gels play an important role in biology and in inspiring biomimetic active materials, due to their ability to change shape, size and create their own morphology; the relevant mechanics behind these changes is driven by…

Soft Condensed Matter · Physics 2023-09-22 Anne Bernheim-Groswasser , Gefen Livne , Paola Nardinocchi , Filippo Recrosi , Luciano Teresi

The actin cytoskeleton of adherent tissue cells often condenses into filament bundles contracted by myosin motors, so-called stress fibers, which play a crucial role in the mechanical interaction of cells with their environment. Stress…

Subcellular Processes · Quantitative Biology 2015-05-27 Achim Besser , Julien Colombelli , Ernst H. K. Stelzer , Ulrich S. Schwarz

The behaviour of elastic structures undergoing large deformations is the result of the competition between confining conditions, self-avoidance and elasticity. This combination of multiple phenomena creates a geometrical frustration that…

Soft Condensed Matter · Physics 2015-05-19 Mokhtar Adda-Bedia , Arezki Boudaoud , Laurent Boué , Stephanie Deboeuf

Cells exhibit qualitatively different behaviors on substrates with different rigidities. The fact that cells are more polarized on the stiffer substrate motivates us to construct a two-dimensional cell with the distribution of focal…

Biological Physics · Physics 2017-07-26 Guangyuan Yu , Jingchen Feng , Haoran Man , Herbert Levine

Using a minimal model of cells or cohesive cell layers as continuum active elastic media, we examine the effect of substrate thickness and stiffness on traction forces exerted by strongly adhering cells. We obtain a simple expression for…

Soft Condensed Matter · Physics 2015-06-04 Shiladitya Banerjee , M. Cristina Marchetti

Based on symmetry consideration of migration and shape deformations, we formulate phenomenologically the dynamics of cell crawling in two dimensions. Forces are introduced to change the cell shape. The shape deformations induce migration of…

Biological Physics · Physics 2016-03-23 Takao Ohta , Mitsusuke Tarama , Masaki Sano

The study of spindle-like cells as nematic liquid crystals has led to remarkable insights in the understanding of tissue organization and morphogenesis. In the characterization of this anomalous liquid crystal material, we focus on the…

Soft Condensed Matter · Physics 2025-07-24 Aniruddh Murali , Prasoon Awasthi , Kirsten Endresen , Arkadiusz Goszczak , Francesca Serra