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Many biological systems are appropriately viewed as passive inclusions immersed in an active bath: from proteins on active membranes to microscopic swimmers confined by boundaries. The non-equilibrium forces exerted by the active bath on…

Soft Condensed Matter · Physics 2020-07-06 Alpha A. Lee , Dominic Vella , John S. Wettlaufer

The response of thermodynamic systems perturbed out of an equilibrium steady-state is described by the reciprocal and the fluctuation-dissipation relations. The so-called fluctuation theorems extended the study of fluctuations far beyond…

Statistical Mechanics · Physics 2020-02-21 Matteo Polettini , Massimiliano Esposito

We use scaling arguments and coarse grained Monte Carlo simulations to study the fluctuation mediated interactions between a pair of adhesion sites of a bilayer membrane and a supporting surface. We find that the potential of mean force is…

Soft Condensed Matter · Physics 2015-05-18 Oded Farago

Microswimmers and active colloids often move in confined systems, including those involving interfaces. Such interfaces, especially at the microscale, may deform in response to the stresses of the flow created by the active particle. We…

Soft Condensed Matter · Physics 2026-01-14 Adam Hitin Bialus , Bhargav Rallabandi , Naomi Oppenheimer

Understanding active matter has led to new perspectives on biophysics and non-equilibrium dynamics. However, the development of numerical tools for simulating active fluids capable of incorporating non-trivial boundaries or inclusions has…

Soft Condensed Matter · Physics 2024-02-01 Timofey Kozhukhov , Benjamin Loewe , Tyler N. Shendruk

We use computer simulations and a simple free energy model to study the response of a bilayer membrane to the application of a negative (compressive) mechanical tension. Such a tension destabilizes the long wavelength undulation modes of…

Soft Condensed Matter · Physics 2015-06-24 Yotam Y. Avital , Oded Farago

Suspensions of Active Brownian Particles (ABP) undergo motility induced phase separation (MIPS) over a wide range of mean density and activity strength [1], even in the absence of an explicit attraction. Negative values of the mechanical…

Soft Condensed Matter · Physics 2022-02-07 Enrique Chacon , Francisco Alarcon , Jorge Ramirez , Pedro Tarazona , Chantal Valeriani

We study the dynamics of a single trapped probe surrounded by self-propelled active particles in two dimensions. In the limit of large size separation, we perform an adiabatic elimination of the small active particles to obtain an effective…

Soft Condensed Matter · Physics 2023-07-21 Ashreya Jayaram , Thomas Speck

In thermodynamic equilibrium, the fluctuation-dissipation theorem links thermal fluctuations and dissipation. Biological systems, however, are driven out of equilibrium by internal processes that produce additional, active fluctuations.…

Soft Condensed Matter · Physics 2026-01-29 Ashot Matevosyan , Frank Jülicher , Ricard Alert

In theory, active control could be used to reduce the unwanted noise reflections from surfaces such as a submarine hull or the walls of an anechoic room. In the recent years, a real-time algorithm has been developed to this effect at the…

Classical Physics · Physics 2009-11-25 Emmanuel Friot , Alexandre Gintz

Theoretical studies of nearly spherical vesicles and microemulsion droplets, that present typical examples for thermally-excited systems that are subject to constraints, are reviewed. We consider the shape fluctuations of such systems…

Soft Condensed Matter · Physics 2021-04-28 Nicholay S. Tonchev

A simple kinetic model of a two-component deformable and reactive bilayer is presented. The two differently shaped components are interconverted by a nonequilibrium reaction, and a phenomenological coupling between local composition and…

Soft Condensed Matter · Physics 2009-11-11 Ramon Reigada , Javier Buceta , Katja Lindenberg

We calculate the mean square amplitude of the shape fluctuation -- an equal-time correlation -- of an almost planar fluid membrane immersed in a near-critical binary fluid mixture. One fluid component is usually preferentially attracted by…

Soft Condensed Matter · Physics 2017-04-26 Youhei Fujitani

Lipid membranes constitute very particular materials: on the one hand, they break very easily under microscopical stretching; on the other hand, they are extremely flexible, presenting deformations even at small scales. Consequently, a…

Soft Condensed Matter · Physics 2010-11-09 Camilla Barbetta

We study the surface fluctuations of a tissue with a dynamics dictated by cell-rearrangement, cell-division, and cell-death processes. Surface fluctuations are calculated in the homeostatic state, where cell division and cell death…

Biological Physics · Physics 2015-12-22 Thomas Risler , Aurélien Peilloux , Jacques Prost

The interaction potential between supported floating bilayers has been determined by grazing incidence specular and off-specular scattering using synchrotron radiation. Our measurements demonstrate that floating bilayers are significantly…

Soft Condensed Matter · Physics 2012-05-21 L. Malaquin , T. Charitat , J. Daillant , S. Lecuyer , G. Fragneto

We experimentally determine the force exerted by a bath of active particles onto a passive probe as a function of its distance to a wall and compare it to the measured averaged density distribution of active particles around the probe.…

Soft Condensed Matter · Physics 2022-08-10 Shuvojit Paul , Ashreya Jayaram , N Narinder , Thomas Speck , Clemens Bechinger

Active tissues exhibit tension fluctuations that are correlated in space and time. We study a minimal overdamped surface model in which such fluctuations enter as a zero-mean, multiplicative modulation of the local surface tension. Although…

Soft Condensed Matter · Physics 2026-03-02 Matteo Ciarchi , Andriy Goychuk , Erwin Frey

The fluctuation pressure of a lipid-bilayer membrane is important for the stability of lamellar phases and the adhesion of membranes to surfaces. In contrast to many theoretical studies, which predict a decrease of the pressure with the…

Soft Condensed Matter · Physics 2013-07-25 Thorsten Auth , Gerhard Gompper

Myosin motor proteins drive vigorous steady-state fluctuations in the actin cytoskeleton of cells. Endogenous embedded semiflexible filaments such as microtubules, or added filaments such as single-walled carbon nanotubes are used as novel…

Biological Physics · Physics 2016-06-21 J. Gladrow , N. Fakhri , F. C. MacKintosh , C. F. Schmidt , C. P. Broedersz