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A striking feature of the collective behavior of spherical microswimmers is that for sufficiently strong self-propulsion they phase-separate into a dense cluster coexisting with a low-density dis- ordered surrounding. Extending our previous…

Soft Condensed Matter · Physics 2016-10-31 Johannes Blaschke , Maurice Maurer , Karthik Menon , Andreas Zöttl , Holger Stark

Active matter has been the object of huge amount of research in recent years for its important fundamental and applicative properties. In this paper we investigate active suspensions of micro-swimmers through direct numerical simulation, so…

Soft Condensed Matter · Physics 2021-02-03 Sergio Chibbaro , Astrid Decoene , Sebastien Martin , Fabien Vergnet

The habitat of microorganisms is typically complex and viscoelastic. A natural question arises: Do polymers in a suspension of active swimmers enhance chaotic motion or promote orientational order? We address this issue by performing…

Soft Condensed Matter · Physics 2025-12-23 Yuan Zhou , Qingzhi Zou , Ignacio Pagonabarraga , Kaihuan Zhang , Kai Qi

An artificial microswimmer drifts in response to spatio-temporal modulations of an activating suspension medium. We consider two competing mechanisms capable of influencing its tactic response: angular fluctuations, which help it explore…

Soft Condensed Matter · Physics 2017-02-15 Alexander Geiseler , Peter Hänggi , Fabio Marchesoni

The acoustofluidic method holds great promise for manipulating microorganisms. When exposed to the steady vortex structures of acoustic streaming flow, these microorganisms exhibit intriguing dynamic behaviors, such as hydrodynamic trapping…

Fluid Dynamics · Physics 2025-04-25 Xuyang Sun , Wenchang Tan , Yi Man

Self-propelled micron-size particles suspended in a fluid, like bacteria or synthetic microswimmers, are strongly non-equilibrium systems where particle motility breaks the microscopic detailed balance, often resulting in large-scale…

Soft Condensed Matter · Physics 2024-03-15 Dóra Bárdfalvy , Viktor Škultéty , Cesare Nardini , Alexander Morozov , Joakim Stenhammar

We study different types of microswimmers moving in channels with varying cross section and thereby interacting hydrodynamically with the channel walls. Starting from the Smoluchowski equation for a dilute suspension, for which interactions…

Soft Condensed Matter · Physics 2017-05-24 Paolo Malgaretti , Holger Stark

Geometric confinements are frequently encountered in the biological world and strongly affect the stability, topology, and transport properties of active suspensions in viscous flow. Based on a far-field analytical model, the…

Microswimmers exhibit more diverse behavior in quasi-two dimensions than in three dimensions. Such behavior remains elusive due to the analytical difficulty of dealing with two parallel solid boundaries. The existence of additional…

Soft Condensed Matter · Physics 2023-01-18 Yuki Takaha , Daiki Nishiguchi

The behavior of fluid interfaces far from equilibrium plays central roles in nature and in industry. Active swimmers trapped at interfaces can alter transport at fluid boundaries with far reaching implications. Swimmers can become trapped…

Soft Condensed Matter · Physics 2024-02-08 Jiayi Deng , Mehdi Molaei , Nicholas G. Chisholm , Kathleen J. Stebe

We consider the tracer diffusion $D_{rr}$ that arises from the run-and-tumble motion of low Reynolds number swimmers, such as bacteria. In unbounded dilute suspensions, where the dipole swimmers move in uncorrelated runs of length…

Soft Condensed Matter · Physics 2015-06-19 Dmitri O. Pushkin , Julia M. Yeomans

Many fascinating properties of biological active matter crucially depend on the capacity of constituting entities to perform directed motion, e.g., molecular motors transporting vesicles inside cells or bacteria searching for food. While…

Statistical Mechanics · Physics 2023-06-21 Pietro Luigi Muzzeddu , Édgar Roldán , Andrea Gambassi , Abhinav Sharma

The trajectories of microswimmers moving in narrow channels of widths comparable to their sizes are significantly altered when they encounter another microswimmer moving in the opposite direction. The consequence of these encounters is a…

The spatiotemporal dynamics in systems of active self-propelled particles is controlled by the propulsion mechanism in combination with various direct interactions, such as steric repulsion, hydrodynamics, and chemical fields. Yet, these…

Soft Condensed Matter · Physics 2024-06-03 Mario Theers , Elmar Westphal , Kai Qi , Roland G. Winkler , Gerhard Gompper

We present a two dimensional model of hydrodynamic interaction between a circular swimmer and a circular post at low Reynolds number, using a point singularity description of the swimming activity. We derive a nonlinear dynamical system…

Fluid Dynamics · Physics 2016-12-09 Dario Papavassiliou , Gareth P Alexander

Suspensions of rear- and front-actuated microswimmers immersed in a fluid, known respectively as ``pushers'' and ``pullers'', display qualitatively different collective behaviours: beyond a characteristic density, pusher suspensions exhibit…

Soft Condensed Matter · Physics 2020-07-08 Dóra Bárdfalvy , Shan Anjum , Cesare Nardini , Alexander Morozov , Joakim Stenhammar

Biological microswimmers often encounter deformable boundaries in physiological conditions; for instance, the viscoelastic walls of reproductive tract during migration of spermatozoa, or host tissue during early bacterial biofilm formation.…

Soft Condensed Matter · Physics 2025-08-07 Smita S. Sontakke , Aneesha Kajampady , Mohd Suhail Rizvi , Ranabir Dey

Hydrodynamic interactions are crucial for determining the cooperative behavior of microswimmers at low Reynolds numbers. Here we provide a comprehensive analysis of the scaling and strength of the interactions in the case of a pair of…

Soft Condensed Matter · Physics 2021-02-11 Sebastian Ziegler , Thomas Scheel , Maxime Hubert , Jens Harting , Ana-Sunčana Smith

Both natural and artificial small-scale swimmers may often self-propel in environments subject to complex geometrical constraints. While most past theoretical work on low-Reynolds number locomotion addressed idealised geometrical…

Fluid Dynamics · Physics 2017-11-16 Alexander Chamolly , Takuji Ishikawa , Eric Lauga

After colliding with a surface, microswimmers reside there during the detention time. They accumulate and may form complex structures such as biofilms. We introduce a general framework to calculate the distribution of detention times using…

Soft Condensed Matter · Physics 2015-07-21 Konstantin Schaar , Andreas Zöttl , Holger Stark