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The collective motion of epithelial cells is a fundamental biological process which plays a significant role in embryogenesis, wound healing and tumor metastasis. While it has been broadly investigated for over a decade both in vivo and in…

We study the stability of the ordered phase of compressible polar flocks against the nucleation of counter-propagating droplets, using a combination of analytical theory, microscopic and hydrodynamic simulations. For discrete-symmetry…

Soft Condensed Matter · Physics 2026-02-20 Omer Granek , Hugues Chaté , Yariv Kafri , Sunghan Ro , Alexandre Solon , Julien Tailleur

Epithelial cell monolayers expand on substrates by forming finger-like protrusions, created by leader cells, in the monolayer boundary. Information transmission and communication between individual entities in the cohesive collective lead…

Biological Physics · Physics 2021-09-06 Sameeksha Rao , Suvakash Dey , Namrata Gundiah

We investigate the segregation of a dense binary mixture of granular particles that only differ in their restitution coefficient. The mixture is vertically vibrated in the presence of gravity. We find a partial segregation of the species,…

Statistical Mechanics · Physics 2009-11-13 R. Brito , H. Enríquez , S. Godoy , R. Soto

We present a comprehensive computational study of the collective behavior emerging from the competition between self-propulsion, excluded volume interactions and velocity-alignment in a two-dimensionnal model of active particles. We…

Soft Condensed Matter · Physics 2018-01-08 Aitor Martín-Gómez , Demian Levis , Albert Díaz-Guilera , Ignacio Pagonabarraga

Recent studies in the collective behavior of active colloids have shown that a global polar order may emerge due to long-ranged chemo-repulsive interactions between them. Here, we report the role of pinning disorder in the flocking…

Soft Condensed Matter · Physics 2026-03-02 Sagarika Adhikary , Arvin Gopal Subramaniam , Rajesh Singh

We show that hydrodynamic theories of polar active matter generically possess inhomogeneous traveling solutions. We introduce a unifying dynamical-system framework to establish the shape of these intrinsically nonlinear patterns, and show…

As the constituents of a flock are polar, one expects a fore-aft asymmetry in their interactions. We show here that the resulting antisymmetric part of the "exchange coupling" between a bird and its neighbours, if large enough, destabilizes…

Statistical Mechanics · Physics 2020-05-06 Lokrshi Prawar Dadhichi , Rahul Chajwa , Ananyo Maitra , Sriram Ramaswamy

In this study, we introduce a minimal model for a collection of polar self-propelled particles (SPPs) on a two-dimensional substrate where each particle has a different ability to interact with its neighbours. The SPPs interact through a…

Statistical Mechanics · Physics 2021-02-02 Jay Prakash Singh , Sudipta Pattanayak , Shradha Mishra

Recent experimental evidence suggests that interactions in flocks of birds do not involve a characteristic length scale. Bird flocks have also been revealed to have an inhomogeneous density distribution, with the density of birds near the…

Quantitative Methods · Quantitative Biology 2019-02-22 Jason M. Lewis , Matthew S. Turner

Flocking, as paradigmatically exemplified by birds, is the coherent collective motion of active agents. As originally conceived, flocking emerges through alignment interactions between the agents. Here, we report that flocking can also…

Soft Condensed Matter · Physics 2024-07-16 Suchismita Das , Matteo Ciarchi , Ziqi Zhou , Jing Yan , Jie Zhang , Ricard Alert

We report flocking in the dry active granular matter of millimeter-sized two-step-tapered rods without an intervening medium. The system undergoes the flocking phase transition at a threshold area fraction ~ 0.12 having high orientational…

Soft Condensed Matter · Physics 2020-06-16 Pradip K. Bera , A. K. Sood

Granular materials are inherently out-of-equilibrium systems due to energy dissipation through inelastic collisions and friction. When driven by mechanical agitation such as vibration, they exhibit rich collective behaviors including…

Soft Condensed Matter · Physics 2026-05-01 Kai Kono , Hiroyuki Ebata , Shio Inagaki

Adding a non-adsorbing polymer to passive colloids induces an attraction between the particles via the `depletion' mechanism. High enough polymer concentrations lead to phase separation. We combine experiments, theory and simulations to…

Soft Condensed Matter · Physics 2016-07-13 J. Schwarz-Linek , C. Valeriani , A. Cacciuto , M. E. Cates , D. Marenduzzo , A. N. Morozov , W. C. K. Poon

We consider a coarse-grained description of a system of self-propelled particles given by hydrodynamic equations for the density and polarization fields. We find that the ordered moving or flocking state of the system is unstable to spatial…

Statistical Mechanics · Physics 2013-05-29 Shradha Mishra , Aparna Baskaran , M. Cristina Marchetti

Steady state dynamics of clustering, long range order, and inelastic collapse are experimentally observed in vertically shaken granular monolayers. At large vibration amplitudes, particle correlations show only short range order like…

Soft Condensed Matter · Physics 2009-10-31 J. S. Olafsen , J. S. Urbach

The effect of structural constriction on molecular mobility is investigated by broadband dielectric spectroscopy (BDS) within three types of molecular arrangements: monomers, oligomers and self-assembled monolayers (SAMs). While disordered…

Collective dynamics in proliferating anisotropic particle systems arise from an interplay between growth, division, and mechanical interactions, often mediated by particle shape. In classical models of prolate, rod-like growth, flow-induced…

Soft Condensed Matter · Physics 2025-12-02 Jonas Isensee , Lukas Hupe , Philip Bittihn

Asymmetric obstacles can be exploited to direct the motion and induce sorting of run-and-tumbling particles. In this work, we show that flocking particles which follow the Vicsek model aligning rules experience a collective trapping in the…

Soft Condensed Matter · Physics 2019-12-02 Raul Martinez , Francisco Alarcon , Juan Luis Aragones , Chantal Valeriani

We investigate the response of colloidal flocks to external fields. We first show that individual colloidal rollers align with external flows as would a classical spin with magnetic fields. Assembling polar active liquids from colloidal…

Soft Condensed Matter · Physics 2018-05-16 Alexandre Morin , Denis Bartolo