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Many microswimmers are able to swim through viscous fluids by employing periodic non-reciprocal deformations of their appendages. Here we use a simple microswimmer model inspired by swimming biflagellates which consists of a spherical cell…

Soft Condensed Matter · Physics 2025-08-22 Sridhar Bulusu , Andreas Zöttl

The green alga {\it Chlamydomonas} swims with synchronized beating of its two flagella, and is experimentally observed to exhibit run-and-tumble behaviour similar to bacteria. Recently we studied a simple hydrodynamic three-sphere model of…

Soft Condensed Matter · Physics 2015-06-15 Rachel R. Bennett , Ramin Golestanian

Run-and-tumble (RT) motion is commonly observed in flagellated microswimmers, arising from synchronous and asynchronous flagellar beating. One such example is a biflagellated alga, called \textit{Chlamydomonas reinhardtii}. Its flagellar…

Soft Condensed Matter · Physics 2025-10-17 Somnath Paramanick , Umashankar Pardhi , Harsh Soni , Nitin Kumar

Many biological microswimmers can modulate their swimming gait to achieve directional control of motility, especially when performing steering towards specific directional cues. This can be achieved without the need for obvious…

The unicellular microalga Chlamydomonas reinhardtii is widely recognized as a premier model living microswimmer for physicists and biophysicists. However, the interest around C. reinhardtii goes beyond its swimming capabilities. In fact,…

Eukaryotic swimming cells such as spermatozoa, algae or protozoa use flagella or cilia to move in viscous fluids. The motion of their flexible appendages in the surrounding fluid induces propulsive forces that balance with the viscous drag…

Soft Condensed Matter · Physics 2023-10-06 Maciej Lisicki , Marcos F. Velho Rodrigues , Eric Lauga

We propose minimal models of one-, two- and three-dimensional micro-swimmers at low Reynolds number with a periodic non-reciprocal motion. These swimmers are either "pushers" or "pullers" of fluid along the swimming axis, or combination of…

Soft Condensed Matter · Physics 2010-04-30 Nobuhiko Watari , Ronald G. Larson

Micro-organisms propel themselves in viscous environments by the periodic, nonreciprocal beating of slender appendages known as flagella. Active materials have been widely exploited to mimic this form of locomotion. However, the realization…

Soft Condensed Matter · Physics 2024-08-06 Ariel Surya Boiardi , Giovanni Noselli

The hydrodynamics of a flagellated microorganism is investigated when swimming close to a planar free-slip surface by means of numerical solu- tions of the Stokes equations obtained via a Boundary Element Method. Depending on the initial…

Fluid Dynamics · Physics 2017-03-31 Daniela Pimponi , Mauro Chinappi , Paolo Gualtieri , Carlo Massimo Casciola

Many biological microorganisms and artificial microswimmers react to external cues of environmental gradients by changing their swimming directions. We study here the behavior of eukarytic flagellated microswimmers in linear viscosity…

Soft Condensed Matter · Physics 2026-05-29 Shubham Anand , Jens Elgeti , Gerhard Gompper

The unicellular biflagellate green alga {\it Chlamydomonas reinhardtii} has been an important model system in biology for decades, and in recent years it has started to attract growing attention also within the biophysics community. Here we…

Cell Behavior · Quantitative Biology 2016-11-23 Raphaël Jeanneret , Matteo Contino , Marco Polin

In a fluid environment, flagellated microswimmers propel themselves by rotating their flagella. The morphology of these flagella significantly influences forward speed, swimming efficiency, and directional stability, which are critical for…

Fluid Dynamics · Physics 2025-06-25 Baopi Liu , Lu Chen , Wenjun Xu

Bacterial swimming is well characterized in uniform liquids at rest. The natural habitat of bacterial swimmers, however, is often dominated by moving fluids and interfaces, resulting in shear flows that may strongly alter bacterial…

We combine a general formulation of microswimmmer equations of motion with a numerical bead-shell model to calculate the hydrodynamic interactions with the fluid, from which the swimming speed, power and efficiency are extracted. From this…

Soft Condensed Matter · Physics 2017-03-07 Bram Bet , Gijs Boosten , Marjolein Dijkstra , René van Roij

Active fluids can develop spontaneous flow instabilities and complex patterns. However, spatio-temporal control of active particles has remained challenging, despite its relevance in biological and applied contexts. Here, we harnessed…

Soft Condensed Matter · Physics 2025-09-23 Isabelle Eisenmann , Marco Vona , Nicolas Desprat , Takuji Ishikawa , Eric Lauga , Raphaël Jeanneret

Most motile bacteria swim in viscous fluids by rotating multiple helical flagellar filaments. These semi-rigid filaments repeatedly join ('bundle') and separate ('unbundle'), resulting in a two-gait random walk-like motion of the cell. In…

Fluid Dynamics · Physics 2020-11-18 Alexander Chamolly , Eric Lauga

A flagellated bacterium navigates fluid environments by rotating its helical flagellar bundle. The wobbling of the bacterial body significantly influences its swimming behavior. To quantify the three underlying motions--precession,…

Soft Condensed Matter · Physics 2026-05-29 Jinglei Hu , Chen Gui , Mingxin Mao , Pu Feng , Yurui Liu , Xiangjun Gong , Gerhard Gompper

The unicellular green algae Chlamydomonas swims with two flagella, which can synchronize their beat. Synchronized beating is required to swim both fast and straight. A long-standing hypothesis proposes that synchronization of flagella…

Cell Behavior · Quantitative Biology 2013-11-26 Veikko Geyer , Frank Jülicher , Jonathon Howard , Benjamin M Friedrich

Suspensions of motile microorganisms can spontaneously form large-scale fluid motion, known as bioconvection, characterized by dense downwelling plumes separated by broad upwelling regions. In this study, we investigate bioconvection in…

Fluid Dynamics · Physics 2025-07-01 S. Gore , I. Gholami , S. O. Ahmed , T. Doskhozhina , S. V. R. Ambadipudi , A. J. Bae , A. Gholami

Microorganisms such as algae and bacteria move in a viscous environment with extremely low Reynolds ($Re$), where the viscous drag dominates the inertial forces. They have adapted to this environment by developing specialized features such…

Robotics · Computer Science 2024-12-10 Nnamdi Chikere , Yasemin Ozkan-Aydin