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We investigate the mechanical origin of polymorphic structures in two-dimensional tubulin assemblies, of which microtubules are the best known example. These structures feature twisted ribbons, flat tubulin sheets, macrotubules, and hoops,…

Soft Condensed Matter · Physics 2022-09-27 Ireth García-Aguilar , Steven Zwaan , Luca Giomi

The Turing instability paradigm is revisited in the context of a multispecies diffusion scheme derived from a self-consistent microscopic formulation. The analysis is developed with reference to the case of two species. These latter share…

Biological Physics · Physics 2012-07-02 Duccio Fanelli , Claudia Cianci , Francesca Di Patti

Growing experimental evidence indicates that topological defects could serve as organizing centers in the morphogenesis of tissues. Here, we provide a quantitative explanation for this phenomenon, rooted in the buckling theory of deformable…

Soft Condensed Matter · Physics 2022-04-21 Ludwig A. Hoffmann , Livio Nicola Carenza , Julia Eckert , Luca Giomi

Thin solids often develop elastic instabilities and subsequently complex, multiscale deformation patterns. Revealing the organizing principles of this spatial complexity has ramifications for our understanding of morphogenetic processes in…

Soft Condensed Matter · Physics 2020-02-28 Oleh Tovkach , Junbo Chen , Monica M. Ripp , Teng Zhang , Joseph D. Paulsen , Benny Davidovitch

Intracellular protein patterns regulate a variety of vital cellular processes such as cell division and motility, which often involve dynamic changes of cell shape. These changes in cell shape may in turn affect the dynamics of…

Pattern Formation and Solitons · Physics 2023-07-19 Laeschkir Würthner , Andriy Goychuk , Erwin Frey

Defects are a ubiquitous feature of ordered media. They have certain universal features, independent of the underlying physical system, reflecting their topological origins. While the topological properties of defects are robust, they…

Soft Condensed Matter · Physics 2021-02-24 Chiqun Zhang , Amit Acharya , Alan C. Newell , Shankar C. Venkataramani

Pattern formation in reaction-diffusion systems where the diffusion terms correspond to a Sturm-Liouville problem are studied. These correspond to a problem where the diffusion coefficient depends on the spatial variable: $\nabla \cdot…

Pattern Formation and Solitons · Physics 2022-11-28 E. A. Calderón-Barreto , J. L. Aragón

We consider a class of singularly perturbed 2-component reaction-diffusion equations which admit bistable traveling front solutions, manifesting as sharp, slow-fast-slow, interfaces between stable homogeneous rest states. In many example…

Analysis of PDEs · Mathematics 2022-12-28 Paul Carter , Arjen Doelman , Kaitlynn Lilly , Erin Obermayer , Shreyas Rao

In this paper, I prove necessary and sufficient conditions for the existence of Turing instabilities in a general system with three interacting species. Turing instabilities describe situations when a stable steady state of a reaction…

Pattern Formation and Solitons · Physics 2024-05-24 Vit Piskovsky

Nature is a blossoming of regular structures, signature of self-organization of the underlying microscopic interacting agents. Turing theory of pattern formation is one of the most studied mechanisms to address such phenomena and has been…

Pattern Formation and Solitons · Physics 2025-10-22 Riccardo Muolo , Lorenzo Giambagli , Hiroya Nakao , Duccio Fanelli , Timoteo Carletti

We consider the classical map proposed previously to be the exact classical analogue of Rydberg Molecules calculated with the approximations relevant to the multi-channel quantum defect theory. The resulting classical map is analyzed at…

Chaotic Dynamics · Physics 2009-10-31 F. Leyvraz , M. Lombardi , T. H. Seligman

Since its foundations, more than one hundred years ago, the field of structural biology has strived to understand and analyze the properties of molecules and their interactions by studying the structure that they take in 3D space. However,…

Biomolecules · Quantitative Biology 2023-02-27 Gabriele Corso

We hereby develop the theory of Turing instability for reaction-diffusion systems defined on complex networks assuming finite propagation. Extending to networked systems the framework introduced by Cattaneo in the 40's, we remove the…

Pattern Formation and Solitons · Physics 2025-10-22 Timoteo Carletti , Riccardo Muolo

In the boundary layer of multicomponent fluid mixtures, the species-specific mass flux in the wall-normal direction is determined by the combination of turbulent-diffusiophoretic diffusion due to composition gradients, and diffusion due to…

Fluid Dynamics · Physics 2018-08-10 Sverre G. Johnsen

Fully developed turbulence is a universal and scale-invariant chaotic state characterized by an energy cascade from large to small scales where the cascade is eventually arrested by dissipation. In this article, we show how to harness these…

Soft Condensed Matter · Physics 2024-04-09 Xander M. de Wit , Michel Fruchart , Tali Khain , Federico Toschi , Vincenzo Vitelli

Shapes of biological membranes are dynamically regulated in living cells. Although membrane shape deformation by proteins at thermal equilibrium has been extensively studied, nonequilibrium dynamics have been much less explored. Recently,…

Biological Physics · Physics 2020-11-19 Naoki Tamemoto , Hiroshi Noguchi

Although the pattern formation on polymer gels has been considered as a result of the mechanical instability due to the volume phase transition, we found a macroscopic surface pattern formation not caused by the mechanical instability. It…

Pattern Formation and Solitons · Physics 2007-05-23 Hiroaki Katsuragi

Hyperuniform materials, characterized by their suppressed density fluctuations and vanishing structure factors as the wave number approaches zero, represent a unique state of matter that straddles the boundary between order and randomness.…

Disordered Systems and Neural Networks · Physics 2024-08-20 Yiwen Tang , Xinzhi Li , Dapeng Bi

Nonlinear reaction-diffusion systems are known to exhibit very many novel spatiotemporal patterns. Fisher equation is a prototype of diffusive equations. In this contribution we investigate the integrability properties of the generalized…

Exactly Solvable and Integrable Systems · Physics 2007-05-23 P. S. Bindu , M. Lakshmanan

Spatial organization of proteins in cells is important for many biological functions. In general, the nonlinear, spatially coupled models for protein-pattern formation are only accessible to numerical simulations, which has limited insight…

Pattern Formation and Solitons · Physics 2021-04-16 Fridtjof Brauns , Jacob Halatek , Erwin Frey