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Engineering synthetic materials that mimic the remarkable complexity of living organisms is a fundamental challenge in science and technology. We study the spatiotemporal patterns that emerge when an active nematicfilm of microtubules and…

Active materials are capable of converting free energy into directional motion, giving rise to striking dynamical phenomena. Developing a general understanding of their structure in relation to the underlying non-equilibrium physics would…

Soft Condensed Matter · Physics 2018-10-17 Nitin Kumar , Rui Zhang , Juan J. de Pablo , Margaret L. Gardel

Biological active matter like the cytoskeleton or tissues are characterized by their ability to transform chemical energy into mechanical stress. In addition, it often exhibits orientational order, which is essential for many cellular and…

Biological Physics · Physics 2025-06-05 Ludovic Dumoulin , Carles Blanch-Mercader , Karsten Kruse

Point-like motile topological defects control the universal dynamics of diverse two-dimensional active nematics ranging from shaken granular rods to cellular monolayers. A comparable understanding in higher dimensions has yet to emerge. We…

Active fluids display spontaneous turbulent-like flows known as active turbulence. Recent work revealed that these flows have universal features, independent of the material properties and of the presence of topological defects. However,…

Soft Condensed Matter · Physics 2026-03-19 Ido Lavi , Ricard Alert , Jean-François Joanny , Jaume Casademunt

Active materials are those in which individual, uncoordinated local stresses drive the material out of equilibrium on a global scale. Examples of such assemblies can be seen across scales from schools of fish to the cellular cytoskeleton…

Soft Condensed Matter · Physics 2022-09-30 Nitin Kumar , Rui Zhang , Steven A. Redford , Juan J. de Pablo , Margaret L. Gardel

Active nematics are fluids in which the components have nematic symmetry and are driven out of equilibrium due to the microscopic generation of an active stress. When the active stress is high, it drives flows in the nematic and can lead to…

Soft Condensed Matter · Physics 2025-11-10 Daniel J. G. Pearce , Berta Martínez-Prat , Jordi Ignés-Mullol , Francesc Sagués

We study a continuum model of an extensile active nematic to show that mesoscale turbulence develops in two stages: (i) ordered regions undergo an intrinsic hydrodynamic instability generating walls, lines of stong bend deformations, (ii)…

Soft Condensed Matter · Physics 2015-06-18 Sumesh P. Thampi , Ramin Golestanian , Julia M. Yeomans

Active matter encompasses different nonequilibrium systems in which individual constituents convert energy into non-conservative forces or motion at the microscale. This review provides an elementary introduction to the role of topology in…

Soft Condensed Matter · Physics 2022-06-28 Suraj Shankar , Anton Souslov , Mark J. Bowick , M. Cristina Marchetti , Vincenzo Vitelli

Active matter is naturally out of equilibrium which results in the emergence of diverse dynamic steady states, including the omnipresent chaotic state known as the active turbulence. However, much less is known how active systems…

Soft Condensed Matter · Physics 2023-04-07 Nika Kralj , Miha Ravnik , Žiga Kos

Topological defects play a prominent role in the physics of two-dimensional materials. When driven out of equilibrium in active nematics, disclinations can acquire spontaneous self-propulsion and drive self-sustained flows upon…

Soft Condensed Matter · Physics 2019-12-06 Suraj Shankar , M. Cristina Marchetti

Our study explores the mixture of polar particles in apolar environment. We employ a coarse-grained approach to model the mixture, where polar particles are in minority. The interaction between polar and apolar components is incorporated…

Soft Condensed Matter · Physics 2025-07-11 Pawan Kumar Mishra , Partha Sarathi Mondal , Pratikshya Jena , Shradha Mishra

Topological defects are at the root of the large-scale organization of liquid crystals. In two-dimensional active nematics, two classes of topological defects of charges $\pm 1/2$ are known to play a major role due to active stresses.…

Soft Condensed Matter · Physics 2022-04-08 Louis Brézin , Thomas Risler , Jean-François Joanny

Active systems, from bacterial suspensions to cellular monolayers, are continuously driven out of equilibrium by local injection of energy from their constituent elements and exhibit turbulent-like and chaotic patterns. Here we demonstrate…

Soft Condensed Matter · Physics 2016-02-04 Amin Doostmohammadi , Michael F. Adamer , Sumesh P. Thampi , Julia M. Yeomans

From the mitotic spindle up to tissues and biofilms, many biological systems behave as active droplets, which often break symmetry and change shape spontaneously. Here, I show that active nematic droplets can experience a fingering…

Soft Condensed Matter · Physics 2022-06-07 Ricard Alert

The study of liquid crystals at equilibrium has led to fundamental insights into the nature of ordered materials, as well as to practical applications such as display technologies. Active nematics are a fundamentally different class of…

Soft Condensed Matter · Physics 2015-08-19 Stephen J. DeCamp , Gabriel S. Redner , Aparna Baskaran , Michael F. Hagan , Zvonimir Dogic

Topological defects in active liquid crystals can be confined by introducing gradients of activity. Here, we examine the dynamical behavior of two defects confined by a sharp gradient of activity that separates an active circular region and…

Soft Condensed Matter · Physics 2021-06-16 Ali Mozaffari , Rui Zhang , Noe Atzin , Juan J. de Pablo

Topological defects in active polar fluids exhibit complex dynamics driven by internally generated stresses, reflecting the deep interplay between topology, flow, and non-equilibrium hydrodynamics. Feedback control offers a powerful means…

Soft Condensed Matter · Physics 2025-07-28 Abhinav Singh , Petros Koumoutsakos

A wide range of living and artificial active matter exists in close contact with substrates and under strong confinement, where in addition to dipolar active stresses, quadrupolar active stresses can become important. Here, we numerically…

Soft Condensed Matter · Physics 2024-04-23 Salik A. Sultan , Mehrana R. Nejad , Amin Doostmohammadi

Topological defects play a central role in the formation and organization of various biological systems. Historically, such nonequilibrium defects have been mainly studied in the context of homogeneous active nematics. Phase-separated…

Soft Condensed Matter · Physics 2023-07-13 Timo Krüger , Ivan Maryshev , Erwin Frey
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