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Large deformations play a central role in the shape transformations of slender active and biological structures. A classical example is the eversion of the Volvox embryo, which demonstrates the need for shell theories that can describe…

Soft Condensed Matter · Physics 2026-03-18 Matteo Taffetani , Matteo Pezzulla

We propose a simple mathematical model to describe the mechanical relaxation of cells within a curved epithelial tissue layer represented by an arbitrary curve in two-dimensional space. This model generalises previous one-dimensional models…

Cellular Automata and Lattice Gases · Physics 2025-01-09 Pascal R. Buenzli , Shahak Kuba , Ryan J. Murphy , Matthew J. Simpson

A body immersed in a nematic liquid crystal disturbs the fluid's preferred molecular configuration and increases its stored elastic energy. In an active nematic, the fluid components also generate a stress in the bulk fluid. By introducing…

Soft Condensed Matter · Physics 2025-06-17 Thomas G. J. Chandler , Saverio E. Spagnolie

We formulate a phenomenological elasto-plastic theory to describe a solid undergoing a structural transition from a square (p4mm) to an oblique (p2) lattice in two dimensions. Within our theory, the components of the strain may be…

Materials Science · Physics 2010-03-18 Arya Paul , Surajit Sengupta , Madan Rao

Vascular and non-vascular cells often form an interconnected network in vitro, similar to the early vascular bed of warm blooded embryos. Our time-lapse recordings show that the network forms by extending sprouts, i.e., multicellular linear…

Cell Behavior · Quantitative Biology 2009-11-13 Andras Szabo , Erica D. Perryn , Andras Czirok

Non-Euclidean plates are a subset of the class of elastic bodies having no stress-free configuration. Such bodies exhibit residual stress when relaxed from all external constraints, and may assume complicated equilibrium shapes even in the…

Soft Condensed Matter · Physics 2009-11-13 Efi Efrati , Eran Sharon , Raz Kupferman

Biological tissues exhibit diverse mechanical and rheological behaviors during morphogenesis. While much is known about tissue phase transitions controlled by structural order and cell mechanics, key questions regarding how tissue-scale…

Soft Condensed Matter · Physics 2026-04-29 Pengyu Yu , Fridtjof Brauns , M. Cristina Marchetti

Stress-stress correlations in crystalline solids with long-range order can be straightforwardly derived using elasticity theory. In contrast, the `emergent elasticity' of amorphous solids, rigid materials characterized by an underlying…

Soft Condensed Matter · Physics 2025-07-04 Jimin Bai , Long-Zhou Huang , Jin Shang , Yun-Jiang Wang , Jie Zhang , Matteo Baggioli

Active gel theory has recently been very successful in describing biologically active materials such as actin filaments or moving bacteria in temporally fixed and simple geometries such as cubes or spheres. Here we develop a computational…

Biological Physics · Physics 2019-07-10 Christian Bächer , Stephan Gekle

I put forward a continuum theory for active nematic gels, defined as fluids or suspensions of orientable rodlike objects endowed with active dynamics, that is based on symmetry arguments and compatibility with thermodynamics. The starting…

Soft Condensed Matter · Physics 2017-11-15 Stefano S Turzi

A new microscopic derivation of the elastic constants of amorphous solids is presented within the framework of nonaffine lattice dynamics, which makes use of a perturbative form of the low-frequency eigenvectors of the dynamical matrix…

Soft Condensed Matter · Physics 2024-08-06 Bingyu Cui , Giancarlo Ruocco , Alessio Zaccone

In the context of finite elasticity, we propose plate models describing the spontaneous bending of nematic elastomer thin films due to variations along the thickness of the nematic order parameters. Reduced energy functionals are deduced…

Analysis of PDEs · Mathematics 2017-02-03 Virginia Agostiniani , Antonio DeSimone

We propose a thermodynamically based approach for constructing effective rate-type constitutive relations describing finite deformations of metamaterials. The effective constitutive relations are formulated as \emph{second-order} in time…

Materials Science · Physics 2025-10-09 Vít Průša , Casey Rodriguez , Ladislav Trnka , Martin Vejvoda

Yield-stress fluids are ubiquitous and encountered in diverse fields ranging from natural muddy flows to industrial applications such as secondary battery electrode slurries and direct ink writing. Despite the proposal of various…

Fluid Dynamics · Physics 2026-05-20 Jehyeok Choi , Ju Min Kim , Kwang Soo Cho

We theoretically explore fluidization of epithelial tissues by active T1 neighbor exchanges. We show that the geometry of cell-cell junctions encodes important information about the local features of the energy landscape, which we support…

Soft Condensed Matter · Physics 2018-08-31 Matej Krajnc , Sabyasachi Dasgupta , Primož Ziherl , Jacques Prost

Surface growth is a crucial component of many natural and artificial processes from cell proliferation to additive manufacturing. In elastic systems surface growth is usually accompanied by the development of geometrical incompatibility…

Soft Condensed Matter · Physics 2019-05-08 Lev Truskinovsky , Giuseppe Zurlo

The rheology of biological tissue is key to processes such as embryo development, wound healing and cancer metastasis. Vertex models of confluent tissue monolayers have uncovered a spontaneous liquid-solid transition tuned by cell shape;…

Soft Condensed Matter · Physics 2023-10-02 Suzanne M. Fielding , James O. Cochran , Junxiang Huang , Dapeng Bi , M. Cristina Marchetti

Developing tissues often maintain mechanical coherence while continuously remodeling through cellular processes such as cell divisions and rearrangements. In this way, they are an example of amorphous solids. In passive amorphous solids,…

Soft Condensed Matter · Physics 2026-05-13 Ali Tahaei , Ahandeep Manna , Marko Popović

In tissue development, wound healing, and cancer invasion, coordinated cell motion arises from active forces produced by the cells. The relationship between force and motion remains unclear, however, because the forces result from a sum of…

Biological Physics · Physics 2025-12-17 Molly McCord , Jacob Notbohm

The rheological properties of biological tissues, though fundamental to many physiological and pathological processes such as embryonic development, wound healing, and tumor progression, remain poorly understood. A recent study showed that…

Soft Condensed Matter · Physics 2026-02-25 Tanmoy Ghosh , Kabir Ramola , Saroj Kumar Nandi