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We investigate how shear influences the dynamics of active particles in dilute to concentrated suspensions. Using apolar active suspensions of squirmers as model systems, we show how their long-time diffusive dynamics can surprisingly slow…

Soft Condensed Matter · Physics 2025-11-14 Zhouyang Ge , John F. Brady , Gwynn J. Elfring

In active matter systems, self-propelled particles can self-organize to undergo collective motion, leading to persistent dynamical behavior out of equilibrium. In cells, cytoskeletal filaments and motor proteins self-organize into complex…

Soft Condensed Matter · Physics 2020-05-19 Jeffrey M. Moore , Tyler N. Thompson , Matthew A. Glaser , Meredith D. Betterton

Geometric confinement plays an important role in the dynamics of natural and synthetic microswimmers from bacterial cells to self-propelled particles in high-throughput microfluidic devices. However, little is known about the effects of…

Fluid Dynamics · Physics 2014-11-13 Alan Cheng Hou Tsang , Eva Kanso

The behaviour of microscopic swimmers has previously been explored near large scale confining geometries and in the presence of very small-scale surface roughness. Here we consider an intermediate case of how a simple microswimmer, the…

Fluid Dynamics · Physics 2021-09-14 Kenta Ishimoto , Eamonn A. Gaffney , David J. Smith

We explore theoretically the complex dynamics and emergent behaviors of spinning spheres immersed in viscous fluid. The particles are coupled to each-other via the fluid in which they are suspended: each particle disturbs the surrounding…

Fluid Dynamics · Physics 2015-09-18 Enkeleida Lushi , Petia M. Vlahovska

We perform Brownian dynamics simulations in two dimensions to study the collective behavior of circle swimmers, which are driven by both, an (effective) translational and rotational self-propulsion, and interact via steric repulsion. We…

Soft Condensed Matter · Physics 2021-06-09 Guo-Jun Liao , Sabine H. L. Klapp

Microswimmers exhibit more diverse behavior in quasi-two dimensions than in three dimensions. Such behavior remains elusive due to the analytical difficulty of dealing with two parallel solid boundaries. The existence of additional…

Soft Condensed Matter · Physics 2023-01-18 Yuki Takaha , Daiki Nishiguchi

Micro-swimmer locomotion in heterogeneous media is increasingly relevant in biological physics due to the prevalence of microorganisms in complex environments. A model for such porous media is the Brinkman fluid which accounts for a sparse…

Fluid Dynamics · Physics 2026-02-05 Francisca Guzman-Lastra , Enkeleida Lushi

Active particles, which interact hydrodynamically, display a remarkable variety of emergent collective phenomena. We use squirmers to model spherical microswimmers and explore the collective behavior of thousands of them under the influence…

Fluid Dynamics · Physics 2017-10-31 Jan-Timm Kuhr , Johannes Blaschke , Felix Rühle , Holger Stark

We investigate the nonequilibrium dynamics of semiflexible polymers driven by motor proteins (MPs) in two-dimensional motility assays under harmonic confinement. Using a coarse-grained agent-based model that incorporates stochastic motor…

Soft Condensed Matter · Physics 2025-09-09 Sandip Roy , Abhishek Chaudhuri , Anil Kumar Dasanna

We use linear stability analysis to show that an isotropic phase of elongated particles with dipolar flow fields can develop nematic order as a result of their activity. We argue that ordering is favoured if the particles are flow-aligning…

Soft Condensed Matter · Physics 2020-03-18 Sreejith Santhosh , Mehrana R. Nejad , Amin Doostmohammadi , Julia M. Yeomans , Sumesh P. Thampi

Semiflexible polymers are ubiquitous in natural and artificial systems, where their intermediate rigidity gives rise to rich structural and dynamical behavior. Confinement plays a central role in these behaviors, as spatial restrictions can…

Soft Condensed Matter · Physics 2026-03-31 Nasir Amiri , Jonathan P. Singer , Xin Yong

We conduct experiments with force-free magnetically-driven rigid helical swimmers in Newtonian and viscoelastic (Boger) fluids. By varying the sizes of the swimmer body and its helical tail, we show that the impact of viscoelasticity…

Fluid Dynamics · Physics 2021-05-05 V. Angeles , F. A. Godinez , J. A. Puente-Velazquez , R. Mendez , E. Lauga , R. Zenit

Populations of swimming microorganisms produce fluid motions that lead to dramatically enhanced diffusion of tracer particles. Using simulations of suspensions of swimming particles in a periodic domain, we capture this effect and show that…

Soft Condensed Matter · Physics 2008-07-22 Patrick T. Underhill , Juan P. Hernández-Ortiz , Michael D. Graham

Scaffold proteins organize cellular processes by bringing signaling molecules into interaction, sometimes by forming large signalosomes. Several of these scaffolds are known to polymerize. Their assemblies should therefore not be understood…

Quantitative Methods · Quantitative Biology 2020-03-06 Andrés Ortiz-Muñoz , Héctor F. Medina-Abarca , Walter Fontana

The current study presents a systematic investigation of the locomotion performance of a swimmer with a wide range of parameter settings. Two-dimensional simulations with the immersed boundary method are employed for the fluid-structure…

Fluid Dynamics · Physics 2023-12-21 Wenkang Wang , Xu Chu

We study the clustering properties of inertial particles in a turbulent viscoelastic fluid. The investigation is carried out by means of direct numerical simulations of turbulence in the Oldroyd-B model. The effects of polymers on the small…

Chaotic Dynamics · Physics 2012-03-22 F. De Lillo , G. Boffetta , S. Musacchio

Viscoelastic flows transition from steady to time-dependent, chaotic dynamics under critical flow conditions, but the implications of geometric disorder for flow stability in these systems are unknown. Utilizing microfluidics, we flow a…

Fluid Dynamics · Physics 2020-04-29 Derek M. Walkama , Nicolas Waisbord , Jeffrey S. Guasto

Biological and artificial microswimmers often have to propel through a variety of environments, ranging from heterogeneous suspending media to strong geometrical confinement. Under confinement, local flow fields generated by microswimmers,…

Soft Condensed Matter · Physics 2024-06-04 Florian A. Overberg , Gerhard Gompper , Dmitry A. Fedosov

Typical bodily and environmental fluids encountered by biological swimmers consist of dissolved macromolecules such as proteins and polymers, often rendering them non Newtonian. To mimic such scenarios, we investigate the motion of swimming…

Soft Condensed Matter · Physics 2023-05-23 Prateek Dwivedi , Atishay Shrivastava , Dipin Pillai , Rahul Mangal