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Self-propelled particles include both self-phoretic synthetic colloids and various micro-organisms. By continually consuming energy, they bypass the laws of equilibrium thermodynamics. These laws enforce the Boltzmann distribution in…

Soft Condensed Matter · Physics 2015-04-15 Michael E. Cates , Julien Tailleur

Multi-component membranes such as ternary mixtures of lipids and cholesterol can exhibit coexistence regions between two liquid phases. When such membranes adhere to a corrugated substrate, the phase separation process strongly depends on…

Soft Condensed Matter · Physics 2009-11-13 Bartosz Rozycki , Thomas R. Weikl , Reinhard Lipowsky

Due to the potential applications in biomedical engineering, it becomes more and more important to understand the process of engulfment and internalization of nanoparticles (NPs) by cell membranes. Despite the fact that the interaction…

Biological Physics · Physics 2023-12-12 Ke Xiao , Jing Li , Rui Ma , Chen-Xu Wu

We computationally studied the phase behavior and dynamics of binary mixtures of active particles, where each 'species' had distinct activities leading to distinct velocities, fast and slow. We obtained phase diagrams demonstrating…

Soft Condensed Matter · Physics 2019-09-09 Thomas Kolb , Daphne Klotsa

Drying of biologically-relevant sessile droplets, including passive systems (like DNA and proteins), as well as active microbial systems comprising bacteria and algae, have garnered considerable attention over the last decades. Distinct…

Soft Condensed Matter · Physics 2023-11-07 Anusuya Pal , Amalesh Gope , Anupam Sengupta

We present a novel Monte Carlo simulation of the phase separation dynamics of a model fluid membrane. Such a phase segregation induces shape changes of the membrane and results in local `budding' under favourable conditions. We present a…

Condensed Matter · Physics 2007-05-23 P. B. Sunil Kumar , Madan Rao

The liquid-liquid phase separation of biomolecules is an important process for intracellular organization. Biomolecular sequence combinatorics leads to a large variety of proteins and nucleic acids which can interact to form a diversity of…

Soft Condensed Matter · Physics 2025-08-27 Aria S. Chaderjian , Sam Wilken , Omar A. Saleh

We study interacting active Brownian particles (ABPs) with a space-dependent swim velocity via simulation and theory. We find that, although an equation of state exists, a mechanical equilibrium does not apply to ABPs in activity…

Soft Condensed Matter · Physics 2024-12-13 Adam Wysocki , Anil K. Dasanna , Heiko Rieger

Bacterial assemblies exhibit rich collective behaviors that control their biological functions, making them a relevant object of study from an active matter physics perspective. Dense bacterial suspensions self-organize into distinct…

Soft Condensed Matter · Physics 2025-04-08 Martin Maliet , Nicolas Fix-Boulier , Ludovic Berthier , Maxime Deforet

The two-dimensional Active Brownian Particles system is meant to be composed of hard disks, that show excluded volume interactions, usually simulated via molecular dynamics using pure repulsive potentials. We show that the softness of the…

The sustenance of life depends on the high degree of organization that prevails through different levels of living organisms, from subcellular structures such as biomolecular complexes and organelles to tissues and organs. The physical…

Soft Condensed Matter · Physics 2019-01-14 Hanieh Falahati , Amir Haji-Akbari

Liquid-liquid phase separation plays a major role in the formation and maintenance of various membrane-less subcellular structures in the cytoplasm and nucleus of cells. Biological condensates contain enhanced concentrations of proteins and…

Soft Condensed Matter · Physics 2023-10-03 Paul C Bressloff

We investigate by time-resolved Synchrotron ultra-small X-ray scattering the dynamics of liquid-liquid phase-separation (LLPS) of gluten protein suspensions following a temperature quench. Samples at a fixed concentration (237 mg/ml) but…

Soft Condensed Matter · Physics 2019-08-27 A. Banc , J. Pincemaille , S. Costanzo , E. Chauveau , M. s. Appavou , M. H. Morel , P. Menut , L. Ramos

Active matter is not only indispensable to our understanding of diverse biological processes, but also provides a fertile ground for discovering novel physics. Many emergent properties impossible for equilibrium systems have been…

Soft Condensed Matter · Physics 2022-05-27 Thibault Bertrand , Chiu Fan Lee

Active matter comprises self-driven units, such as bacteria and synthetic microswimmers, that can spontaneously form complex patterns and assemble into functional microdevices. These processes are possible thanks to the out-of-equilibrium…

Soft Condensed Matter · Physics 2021-11-17 Jens Grauer , Falko Schmidt , Jesus Pineda , Benjamin Midtvedt , Hartmut Löwen , Giovanni Volpe , Benno Liebchen

We study the behaviour of catalytically active droplets in multi-component conserved mixtures affected by noise. Working in the thin interface limit, we analytically determine the state diagram of the system, characterized by multiple…

Soft Condensed Matter · Physics 2025-12-01 Jacopo Romano , Martin Kjøllesdal Johnsrud , Benoît Mahault , Ramin Golestanian

Meso-scale turbulence was originally observed experimentally in various suspensions of swimming bacteria, as well as in the collective motion of active colloids. The corresponding large-scale dynamical patterns were reproduced in a simple…

Soft Condensed Matter · Physics 2021-03-30 Vasco M. Worlitzer , Gil Ariel , Avraham Be'er , Holger Stark , Markus Bär , Sebastian Heidenreich

The ion-activated patchy particle model is an important theoretical framework to investigate the phase behaviour of globular proteins in the presence of multivalent ions. In this work, we study and highlight the influence of patch…

Soft Condensed Matter · Physics 2025-11-13 Furio Surfaro , Peixuan Liang , Hadra Banks , Fajun Zhang , Frank Schreiber , Martin Oettel

Depinning and nonequilibrium transitions within sliding states in systems driven over quenched disorder arise across a wide spectrum of size scales ranging from atomic friction at the nanoscale, flux motion in type-II superconductors at the…

Soft Condensed Matter · Physics 2017-03-22 Cs. Sándor , A. Libál , C. Reichhardt , C. J. Olson Reichhardt

With exquisite precision and reproducibility, cells orchestrate the cooperative action of thousands of nanometer-sized molecular motors to carry out mechanical tasks at much larger length scales, such as cell motility, division and…

Soft Condensed Matter · Physics 2013-01-08 Tim Sanchez , Daniel T. N. Chen , Stephen J. DeCamp , Michael Heymann , Zvonimir Dogic