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Under partial confinement, the motion of colloidal particles is restricted to a plane but their dynamics is influenced by hydrodynamic interactions mediated by the unconfined, three--dimensional flow of the embedding fluid. We demonstrate…

Soft Condensed Matter · Physics 2013-05-17 J. Bleibel , A. Dominguez , F. Günther , J. Harting , M. Oettel

Controlling the phases of matter is a challenge that spans from condensed materials to biological systems. Here, by imposing a geometric boundary condition, we study controlled collective motion of Escherichia coli bacteria. A circular…

Soft Condensed Matter · Physics 2017-09-01 Kazusa Beppu , Ziane Izri , Jun Gohya , Kanta Eto , Masatoshi Ichikawa , Yusuke T. Maeda

Interactions between swimming cells and surfaces are essential to many microbiological processes, from bacterial biofilm formation to human fertilization. However, in spite of their fundamental importance, relatively little is known about…

Biological Physics · Physics 2013-02-04 Vasily Kantsler , Jörn Dunkel , Marco Polin , Raymond E. Goldstein

Recent studies aimed at investigating artificial analogs of bacterial colonies have shown that low-density suspensions of self-propelled particles confined in two dimensions can assemble into finite aggregates that merge and split, but have…

Soft Condensed Matter · Physics 2015-06-17 B. M. Mognetti , A. Šarić , S. Angioletti-Uberti , A. Cacciuto , C. Valeriani , D. Frenkel

Collective cell flows are a hallmark of tissue dynamics in development, wound healing, and various diseases. Here, we perform experiments on epithelial MDCK cell monolayers, over tens of hours without jamming, on millimeter-scale…

Tissues and Organs · Quantitative Biology 2025-11-27 Guillaume Duprez , Mélina Durande , François Graner , Hélène Delanoë-Ayari

When an ensemble of particles interact hydrodynamically, they generically display large-scale transient structures such as swirls in sedimenting particles [1], or colloidal strings in sheared suspensions [2]. Understanding these…

Soft Condensed Matter · Physics 2013-01-24 Nicolas Desreumaux , Jean-Baptiste Caussin , Raphael Jeanneret , Eric Lauga , Denis Bartolo

How thousands of microtubules and molecular motors self-organize into spindles remains poorly understood. By combining static, nanometer-resolution, large-scale electron tomography reconstructions and dynamic, optical-resolution, polarized…

Many marine invertebrates have larval stages covered in linear arrays of beating cilia, which propel the animal while simultaneously entraining planktonic prey. These bands are strongly conserved across taxa spanning four major superphyla,…

Fluid Dynamics · Physics 2017-02-14 William Gilpin , Vivek N. Prakash , Manu Prakash

While the collective motion of active particles has been studied extensively, effective strategies to navigate particle swarms without external guidance remain elusive. We introduce a method to control the trajectories of two-dimensional…

Soft Condensed Matter · Physics 2023-08-28 Sophie Y. Lee , Philipp W. A. Schönhöfer , Sharon C. Glotzer

Living organisms employ diverse strategies to navigate confined environments. Inspired by translocation observations on California blackworms (\textit{Lumbriculus variegatus}), we combine biological experiments and active-polymer…

Soft Condensed Matter · Physics 2025-10-21 K. R. Prathyusha , Paulami Sarkar , Justin Xu , Saad Bhamla

One of the major challenges in modern robotics is controlling micromanipulation by active and adaptive materials. In the respiratory system, such actuation enables pathogen clearance by means of motile cilia. While various types of…

Hypothesis: New dynamic phenomena can be observed in evaporating free liquid films from colloidal solutions with bimodal particle size distribution. Such distributions are formed in a natural way in mixed (slightly turbid) solutions of…

Soft Condensed Matter · Physics 2020-05-21 E. S. Basheva , P. A. Kralchevsky , K. D. Danov , R. D. Stanimirova , N. Shaw , J. T. Petkov

Vortex rings are critical for thrust production underwater. In the ocean, self-propelled mesozooplankton generate vortices while swimming within a weakly stratified fluid. While large-scale biogenic transport has been observed during…

Fluid Dynamics · Physics 2023-04-12 Yunxing Su , Monica M. Wilhelmus , Roberto Zenit

We study active agents embedded in bulk or in confinement explicitly considering hydrodynamics and simulating the swimmers via an implementation inspired by the squirmer model. We develop a Dissipative Particle Dynamics scheme for the…

The ocean is teeming with a myriad of mm-sized invertebrate planktonic larvae, which thrive in a viscous fluid environment. Many of them rely on ciliary beating to generate fluid flows for locomotion and feeding. Their larval forms, local…

Hydrodynamics and confinement dominate bacterial mobility near solid or air-water boundaries, causing flagellated bacteria to move in circular trajectories. This phenomenon results from the counter-rotation between the bacterial body and…

Biological Physics · Physics 2018-10-09 George Araujo , Weijie Chen , Sridhar Mani , Jay X. Tang

We analyze the dynamics of small particles vertically confined, by means of a linear restoring force, to move within a horizontal fluid slab in a three-dimensional (3D) homogeneous isotropic turbulent velocity field. The model that we…

Biological and artificial microswimmers often self-propel in external flows of vortical nature; relevant examples include algae in small-scale ocean eddies, spermatozoa in uterine peristaltic flows and bacteria in microfluidic devices. A…

Biological Physics · Physics 2022-11-14 Ivan Tanasijevic , Eric Lauga

Ciliated tissues such as in the mammalian lungs, brains, and reproductive tracts, are specialized to pump fluid. They generate flows by the collective activity of hundreds of thousands of individual cilia that beat in a striking metachronal…

Soft Condensed Matter · Physics 2022-11-30 Anup Kanale , Feng Ling , Hanliang Guo , Sebastian Fuerthauer , Eva Kanso

Understanding mixing and transport of passive scalars in active fluids is important to many natural (e.g. algal blooms) and industrial (e.g. biofuel, vaccine production) processes. Here, we study the mixing of a passive scalar (dye) in…