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Synchronisation and pattern formation have been intensely addressed for systems evolving on static networks. Extending the study to include the inherent ability of the network to adjust over time proved cumbersome and led to conclusions…

Statistical Mechanics · Physics 2022-05-25 Timoteo Carletti , Duccio Fanelli

Pattern formation in reaction-diffusion systems where the diffusion terms correspond to a Sturm-Liouville problem are studied. These correspond to a problem where the diffusion coefficient depends on the spatial variable: $\nabla \cdot…

Pattern Formation and Solitons · Physics 2022-11-28 E. A. Calderón-Barreto , J. L. Aragón

Analytically tracking patterns emerging from a small amplitude Turing instability to large amplitude remains a challenge as no general theory exists. In this paper, we consider a three component reaction-diffusion system with one of its…

Dynamical Systems · Mathematics 2023-11-06 Christopher Brown , Gianne Derks , Peter van Heijster , David J. B. Lloyd

Reaction-diffusion processes on networked systems have received mounting attention in the past two decades, and the corresponding theory of network dynamics has been continuously enriched with the advancement of network science. Recently,…

Pattern Formation and Solitons · Physics 2025-04-30 Junyuan Shi , Linhe Zhu

Turing's theory of pattern formation is a universal model for self-organization, applicable to many systems in physics, chemistry and biology. Essential properties of a Turing system, such as the conditions for the existence of patterns and…

Adaptation and Self-Organizing Systems · Physics 2018-08-02 Xavier Diego , Luciano Marcon , Patrick Müller , James Sharpe

In this work we investigate the process of pattern formation in a two dimensional domain for a reaction-diffusion system with nonlinear diffusion terms and the competitive Lotka-Volterra kinetics. The linear stability analysis shows that…

Pattern Formation and Solitons · Physics 2014-03-03 G. Gambino , M. C. Lombardo , M. Sammartino

It is well known that simple reaction-diffusion systems can display very rich pattern formation behavior. Here we have studied two examples of such systems in three dimensions. First we investigate the morphology and stability of a generic…

Statistical Mechanics · Physics 2009-11-07 Teemu Leppanen , Mikko Karttunen , Kimmo Kaski , Rafael A. Barrio , Limei Zhang

In this paper, we analyse the dynamics of a pattern-forming system close to simultaneous Turing and Turing--Hopf instabilities, which have a 1:1 spatial resonance, that is, they have the same critical wave number. For this, we consider a…

Analysis of PDEs · Mathematics 2025-09-01 Bastian Hilder , Christian Kuehn

Subcritical Turing bifurcations of reaction-diffusion systems in large domains lead to spontaneous onset of well-developed localised patterns via the homoclinic snaking mechanism. This phenomenon is shown to occur naturally when balancing…

Pattern Formation and Solitons · Physics 2025-06-10 Víctor Breña-Medina , Alan Champneys

We analyzed conditions for Hopf and Turing instabilities to occur in two-component fractional reaction-diffusion systems. We showed that the eigenvalue spectrum and fractional derivative order mainly determine the type of instability and…

Adaptation and Self-Organizing Systems · Physics 2009-12-09 B. Y. Datsko , V. V. Gafiychuk

The process of stochastic Turing instability on a network is discussed for a specific case study, the stochastic Brusselator model. The system is shown to spontaneously differentiate into activator-rich and activator-poor nodes, outside the…

Statistical Mechanics · Physics 2015-06-04 Malbor Asslani , Francesca Di Patti , Duccio Fanelli

GTPase molecules are important regulators in cells that continuously run through an activation/deactivation and membrane-attachment/membrane-detachment cycle. Activated GTPase is able to localize in parts of the membranes and to induce cell…

Analysis of PDEs · Mathematics 2011-12-08 Andreas Rätz , Matthias Röger

Turing (or double-diffusive) instabilities describe pattern formation in reaction-diffusion systems, and were proposed in 1952 as a potential mechanism behind pattern formation in nature, such as leopard spots and zebra stripes. Because the…

Materials Science · Physics 2020-04-29 M. W. Noble , M. R. Tonks , S. P. Fitzgerald

We explain the principles of gene expression pattern stabilization in systems of interacting, diffusible morphogens, with dynamically established source regions. Using a reaction-diffusion model with step-function production term, we…

Biological Physics · Physics 2023-03-02 M. Majka , R. D. J. G. Ho , M. Zagorski

Spatial distribution of the human population is distinctly heterogeneous, e.g. showing significant difference in the population density between urban and rural areas. In the historical perspective, i.e. on the timescale of centuries, the…

Adaptation and Self-Organizing Systems · Physics 2022-08-30 Anna Zincenko , Sergei Petrovskii , Vitaly Volpert

This paper is concerned with stochastic reaction-diffusion kinetics governed by the reaction-diffusion master equation. Specifically, the primary goal of this paper is to provide a mechanistic basis of Turing pattern formation that is…

Quantitative Methods · Quantitative Biology 2015-04-23 Yutaka Hori , Shinji Hara

We hereby develop the theory of Turing instability for reaction-diffusion systems defined on complex networks assuming finite propagation. Extending to networked systems the framework introduced by Cattaneo in the 40's, we remove the…

Pattern Formation and Solitons · Physics 2025-10-22 Timoteo Carletti , Riccardo Muolo

In this paper, I prove necessary and sufficient conditions for the existence of Turing instabilities in a general system with three interacting species. Turing instabilities describe situations when a stable steady state of a reaction…

Pattern Formation and Solitons · Physics 2024-05-24 Vit Piskovsky

A class of systems is considered, where immobile species associated to distinct patches, the nodes of a network, interact both locally and at a long-range, as specified by an (interaction) adjacency matrix. Non local interactions are…

Pattern Formation and Solitons · Physics 2020-06-30 Giulia Cencetti , Federico Battiston , Timoteo Carletti , Duccio Fanelli

The rheological properties of biological tissues are core to processes such as cancer metastasis, wound healing and embryo development. The emergence of tissue and organ structures during morphogenesis requires the precise formation of…

Soft Condensed Matter · Physics 2026-03-10 Aidan J. Nicholas , Suzanne M. Fielding