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The evolutionary balance between innate and learned behaviors is highly intricate, and different organisms have found different solutions to this problem. We hypothesize that the emergence and exact form of learning behaviors is naturally…

Neurons and Cognition · Quantitative Biology 2023-03-14 Emmanouil Giannakakis , Sina Khajehabdollahi , Anna Levina

The nonlinear mechanics of a flexible elastic rod constrained at its edges by a pair of sliding sleeves is analyzed. The planar equilibrium configurations of this variable-length elastica are found to have shape defined only by the…

Soft Condensed Matter · Physics 2024-09-19 Alessandro Cazzolli , Francesco Dal Corso

The accumulation of self-propelled particles on repulsive barriers is a widely observed feature in active matter. Despite being implicated in a broad range of biological processes, from biofilm formation to cytoskeletal movement, wetting of…

Statistical Mechanics · Physics 2025-12-12 Noah Grodzinski , Michael E. Cates , Robert L. Jack

The intermittent transition between slow growth and rapid shrinkage in polymeric assemblies is termed dynamic instability, a feature observed in a variety of biochemically distinct assemblies including microtubules, actin and their…

Soft Condensed Matter · Physics 2015-05-28 Stefano Zapperi , L. Mahadevan

Living soft tissues appear to promote the development and maintenance of a preferred mechanical state within a defined tolerance around a so-called set-point. This phenomenon is often referred to as mechanical homeostasis. In contradiction…

Computational Engineering, Finance, and Science · Computer Science 2021-09-23 Jonas F. Eichinger , Maximilian J. Grill , Iman Davoodi Kermani , Roland C. Aydin , Wolfgang A. Wall , Jay D. Humphrey , Christian J. Cyron

The internal dynamics of active gels, both in artificial (in-vitro) model systems and inside the cytoskeleton of living cells, has been extensively studied by experiments of recent years. These dynamics are probed using tracer particles…

Soft Condensed Matter · Physics 2015-07-28 E. Ben Isaac , É. Fodor , P. Visco , F. van Wijland , N. S. Gov

The elastic properties of a material with spherical voids of equal volume are analysed using a new model, with particular attention paid to the hexagonal close-packed and the face-centred cubic arrangement of voids. Void fractions well…

Materials Science · Physics 2016-11-11 Sascha Heitkam , Wiebke Drenckhan , Denis Weaire , Jochen Froehlich

In these Lecture Notes we aim at clarifying how soft matter physics, and herein notably statistical mechanics and fluid mechanics, can be engaged to understand and manipulate non-equilibrium systems consisting of numerous (microscopic)…

Soft Condensed Matter · Physics 2025-05-27 Jean-François Joanny , Joseph O. Indekeu

Soft composite solids are made of inclusions dispersed within soft matrices. They are ubiquitous in nature and form the basis of many biological tissues. In the field of materials science, synthetic soft composites are promising candidates…

Soft Condensed Matter · Physics 2024-06-14 Yiqiu Zhao , Haitao Hu , Yulu Huang , Hanqing Liu , Caishan Yan , Chang Xu , Rui Zhang , Yifan Wang , Qin Xu

Active materials are capable of converting free energy into mechanical work to produce autonomous motion, and exhibit striking collective dynamics that biology relies on for essential functions. Controlling those dynamics and transport in…

With increasing emphasis on the study of active solids, the features of these classes of nonequilibrium systems and materials beyond their mere existence shift into focus. One concept of active solids addresses them as active,…

Soft Condensed Matter · Physics 2025-02-10 Yutaka Kinoshita , Nariya Uchida , Andreas M. Menzel

The dynamics of active viscoelastic surfaces plays an important role in biological systems. One prominent example is the actin cortex, a thin bio-polymer sheet underneath the outer membrane of biological cells which combines active…

A huge number of technological and biological systems involves the lubricated contact between rough surfaces of soft solids in relative accelerated motion. Examples include dynamical rubber seals and the human joints. In this study we…

Soft Condensed Matter · Physics 2017-02-09 M. Scaraggi , L. Dorogin , J. Angerhausen , H. Murrenhoff , B. N. J. Persson

Active colloids, also known as artificial microswimmers, are self-propelled micro and nanoparticles that convert uniform sources of fuel (e.g. chemical) or uniform external driving fields (e.g. magnetic or electric) into directed motion by…

Soft Condensed Matter · Physics 2017-01-30 Songbo Ni , Emanuele Marini , Ivo Buttinoni , Heiko Wolf , Lucio Isa

Phase separating systems that are maintained away from thermodynamic equilibrium via molecular processes represent a class of active systems, which we call active emulsions. These systems are driven by external energy input for example…

Soft Condensed Matter · Physics 2019-06-07 Christoph A. Weber , David Zwicker , Frank Jülicher , Chiu Fan Lee

Passive elastic elements can contribute to stability, energetic efficiency, and impact absorption in both biological and robotic systems. They also add dynamical complexity which makes them more challenging to model and control. The impact…

Robotics · Computer Science 2019-09-30 Ali Marjaninejad , Jie Tan , Francisco J. Valero-Cuevas

A continuum description is essential for understanding a variety of collective phenomena in active matter. However, building quantitative continuum models of active matter from first principles can be extremely challenging due to both the…

Soft Condensed Matter · Physics 2022-10-19 Matthew Golden , Roman Grigoriev , Jyothishraj Nambisan , Alberto Fernandez-Nieves

Active fluid droplets are a class of soft materials exhibiting autonomous motion sustained by an energy supply. Such systems have been shown to capture motility regimes typical of biological cells and are ideal candidates as building-block…

Soft Condensed Matter · Physics 2023-03-01 A. Tiribocchi , M. Durve , M. Lauricella , A. Montessori , D. Marenduzzo , S. Succi

Elasticity of soft materials can be greatly influenced by the presence of air bubbles. Such a capillary effect is expected for a wide range of materials, from polymer gels to concentrated emulsions and colloidal suspensions. Whereas…

Soft Condensed Matter · Physics 2017-04-11 François Gorlier , Yacine Khidas , Olivier Pitois

Steady structures originating from dynamic self-assembly have begun to show their advantages in new generation materials, and pose challenges to equilibrium self-assembly. In view of the important role of confinement in self-assembly, here,…

Computational Physics · Physics 2019-08-06 Rui-fen Zhang , Chun-lai Ren , Jia-wei Feng , Yu-qiang Ma