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Collective cell migration is a key driver of embryonic development, wound healing, and some types of cancer invasion. Here we provide a physical perspective of the mechanisms underlying collective cell migration. We begin with a catalogue…

Biological Physics · Physics 2019-10-08 Ricard Alert , Xavier Trepat

Inspired by the wake-surfing nature of animals, this study aims to understand the aerodynamic force variation on a wing surfing in an unsteady 2-D wake. Wind tunnel experiments were conducted using Particle Image Velocimetry (PIV) and force…

Fluid Dynamics · Physics 2025-12-11 Siyang Hao , Kenneth Breuer

We investigate the dynamics of $N$ point vortices in the plane, in the limit of large $N$. We consider {\em relative equilibria}, which are rigidly rotating lattice-like configurations of vortices. These configurations were observed in…

Quantum Gases · Physics 2014-02-04 Yuxin Chen , Theodore Kolokolnikov , Daniel Zhirov

Stretching and retracting wingspan has been widely observed in the flight of birds and bats, and its effects on the aerodynamic performance particularly lift generation are intriguing. The rectangular flat-plate flapping wing with a…

Fluid Dynamics · Physics 2021-05-12 Shizhao Wang , Xing Zhang , Guowei He , Tianshu Liu

Flocking is a paradigmatic example of collective animal behaviour, where decentralized interaction rules give rise to a globally ordered state. In the emergence of order out of self-organization we find similarities between biological…

Populations and Evolution · Quantitative Biology 2013-02-14 Andrea Cavagna , Silvio M. Duarte Queiros , Irene Giardina , Fabio Stefanini , Massimiliano Viale

Collective motion is ubiquitous in nature; groups of animals, such as fish, birds, and ungulates appear to move as a whole, exhibiting a rich behavioral repertoire that ranges from directed movement to milling to disordered swarming.…

Adaptation and Self-Organizing Systems · Physics 2024-05-15 Conor Heins , Beren Millidge , Lancelot da Costa , Richard Mann , Karl Friston , Iain Couzin

From the formation of animal flocks to the emergence of coordinate motion in bacterial swarms, at all scales populations of motile organisms display coherent collective motion. This consistent behavior strongly contrasts with the difference…

Soft Condensed Matter · Physics 2013-11-11 Antoine Bricard , Jean-Baptiste Caussin , Nicolas Desreumaux , Olivier Dauchot , Denis Bartolo

Efficient collective response to external perturbations is one of the most striking abilities of a biological system. Signal propagation through the group is an important condition for the imple- mentation of such a response. Information…

Statistical Mechanics · Physics 2018-11-14 Andrea Cavagna , Daniele Conti , Irene Giardina , Tomas S. Grigera

Flocking, as paradigmatically exemplified by birds, is the coherent collective motion of active agents. As originally conceived, flocking emerges through alignment interactions between the agents. Here, we report that flocking can also…

Soft Condensed Matter · Physics 2024-07-16 Suchismita Das , Matteo Ciarchi , Ziqi Zhou , Jing Yan , Jie Zhang , Ricard Alert

The collective behavior of active semiflexible filaments is studied with a model of tangentially driven self-propelled worm-like chains. The combination of excluded-volume interactions and self-propulsion leads to several distinct dynamic…

Biological Physics · Physics 2024-06-03 Ozer Duman , Rolf E. Isele-Holder , Jens Elgeti , Gerhard Gompper

Linear stability analysis of an elastically anchored wing in a uniform flow is investigated both analytically and numerically. The analytical formulation explicitly takes into account the effect of the wake on the wing by means of…

Fluid Dynamics · Physics 2012-02-27 A. Orchini , A. Mazzino , J. Guerrero , R. Festa , C. Boragno

We experimentally study a scallop-like swimmer with reciprocally flapping wings in a nearly frictionless, cohesive granular medium consisting of hydrogel spheres. Significant locomotion is found when the swimmer's flapping frequency matches…

Soft Condensed Matter · Physics 2026-04-29 Hongyi Xiao , Jing Wang , Achim Sack , Ralf Stannarius , Thorsten Pöschel

We generalize the Vicsek model to describe the collective behaviour of polar circle swimmers with local alignment interactions. While the phase transition leading to collective motion in 2D (flocking) occurs at the same interaction to noise…

Statistical Mechanics · Physics 2017-08-09 Benno Liebchen , Demian Levis

This expository review is devoted to fish swimming and bird/insect flight. (i) The simple waving motion of an elongated flexible ribbon plate of constant width, immersed in a fluid at rest, propagating a wave distally down the plate to swim…

Mathematical Physics · Physics 2010-07-30 Theodore Yaotsu Wu

We present and analyze a theoretical model for the dynamics and interactions of "capillary surfers," which are millimetric objects that self-propel while floating at the interface of a vibrating fluid bath. In our companion paper [1], we…

Fluid Dynamics · Physics 2024-11-25 Anand U. Oza , Giuseppe Pucci , Ian Ho , Daniel M. Harris

We explore a simplified class of models we call swarms, which are inspired by the collective behavior of social insects. We perform a mean-field stability analysis and perform numerical simulations of the model. Several interesting types of…

adap-org · Physics 2009-10-28 Erik M. Rauch , Mark M. Millonas , Dante R. Chialvo

This study delves into the aerodynamic behavior of a highly flexible NACA 0012 aerofoil, drawing inspiration from avian feathers to handle a gust. We first examined unsteady flow on a rigid wing both experimentally and numerically and then…

In order to keep their cohesiveness during locomotion gregarious animals must make collective decisions. Many species boast complex societies with multiple levels of communities. A common case is when two dominant levels exist, one…

Biological Physics · Physics 2017-04-03 Bence Ferdinandy , Katalin Ozogány , Tamás Vicsek

Here we show that coupling to curvature has profound effects on collective motion in active systems, leading to patterns not observed in flat space. Biological examples of such active motion in curved environments are numerous: curvature…

Soft Condensed Matter · Physics 2015-03-05 Rastko Sknepnek , Silke Henkes

We explore theoretically the complex dynamics and emergent behaviors of spinning spheres immersed in viscous fluid. The particles are coupled to each-other via the fluid in which they are suspended: each particle disturbs the surrounding…

Fluid Dynamics · Physics 2015-09-18 Enkeleida Lushi , Petia M. Vlahovska