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Active stresses in biological cells and tissues drive many developmental processes. However, increasing experimental evidence suggests that additional mechanical interactions with surrounding material can play a crucial role in guiding…

Soft Condensed Matter · Physics 2026-03-10 Douglas MacMyn Brown , Alexander Mietke

In equilibrium, disorder conspires with topological defects to redefine the ordered states of matter in systems as diverse as crystals, superconductors and liquid crystals. Far from equilibrium, however, the consequences of quenched…

Soft Condensed Matter · Physics 2021-03-09 Amélie Chardac , Suraj Shankar , M. Cristina Marchetti , Denis Bartolo

Active forces occurring within cells can drive crucial biological processes that involve spontaneous organization and shape change, such as cell division. Motivated by recent in vitro experiments of nematic droplets of cytoskeletal…

Soft Condensed Matter · Physics 2021-02-16 Fabian Jan Schwarzendahl , Pierre Ronceray , Kimberly L. Weirich , Kinjal Dasbiswas

Water ice's remarkable properties make it an important material across a range of disciplines. The combination of covalent and hydrogen bonds form a long-range lattice of oxygens, which hosts a disordered yet correlated hydrogen network. We…

We analyze the behavior of a suspension of active polar particles under shear. In the absence of external forces, orientationally ordered active particles are known to exhibit a transition to a state of non-uniform polarization and…

Soft Condensed Matter · Physics 2010-05-07 Luca Giomi , M. Cristina Marchetti , Tanniemola B. Liverpool

Cellular rearrangements, as primary sources of tissue fluidization, facilitate topological transitions during tissue morphogenesis. We study the role of intrinsic cell properties such as cell polarity and cell-cell adhesion in shaping…

Biological Physics · Physics 2024-03-11 Richard D. J. G. Ho , Stig Ove Bøe , Dag Kristian Dysthe , Luiza Angheluta

We study the spatio-temporal dynamics of a model of polar active fluid in two dimensions. The system exhibits a transition from an isotropic to a polarized state as a function of density. The uniform polarized state is, however, unstable…

Soft Condensed Matter · Physics 2011-12-08 Luca Giomi , M. Cristina Marchetti

Crystallography typically studies collections of point particles whose interaction forces are the gradient of a potential. Lifting this assumption generically gives rise in the continuum limit to a form of elasticity with additional moduli…

Soft Condensed Matter · Physics 2021-12-30 Lara Braverman , Colin Scheibner , Bryan VanSaders , Vincenzo Vitelli

We study the effects of inertia in dense suspensions of polar swimmers. The hydrodynamic velocity field and the polar order parameter field describe the dynamics of the suspension. We show that a dimensionless parameter $R$ (ratio of the…

Soft Condensed Matter · Physics 2024-08-14 Navdeep Rana , Rayan Chatterjee , Sunghan Ro , Dov Levine , Sriram Ramaswamy , Prasad Perlekar

A survey is presented of the dynamic features of non-itinerant off-center defects in crystals, such as rotation-like reorientation of isolated species by either impurity or host ions. The occurrence of off-center displacements in…

Chemical Physics · Physics 2008-12-12 Mladen Georgiev

Using novel micro-printing techniques, we develop a versatile experimental setup that allows us to study how lateral confinement tames the active flows and defect properties of the microtubule/kinesin active nematic system. We demonstrate…

Exploring structural order in disordered systems including liquids and glasses is an intriguing but challenging issue in condensed matter physics. Here we construct a new parameter based on the angular distribution function of particles and…

Soft Condensed Matter · Physics 2018-06-11 Xiunan Yang , Wei-Hua Wang , Ke Chen

The transition from complex-periodic to chaotic behavior is investigated in oscillatory media supporting spiral waves. We find turbulent regimes characterized by the spontaneous nucleation, proliferation and erratic motion of…

chao-dyn · Physics 2009-10-31 Andrei Goryachev , Hugues Chate' , Raymond Kapral

Hydrodynamic interactions can generate rich emergent structures in active matter systems. Using large-scale hydrodynamic simulations, we demonstrate that hydrodynamic coupling alone can drive spontaneous self-organization across a hierarchy…

Soft Condensed Matter · Physics 2025-11-17 Zaiyi Shen , Leilei Wang , Shishuang Zhang , Chenlu Li , Kaili Xie , Xu Zheng , Juho S. Lintuvuori

This work introduces a new theoretical model for active matter ("complementary-spins" or c-spins), exploring the interplay of positional and orientational order in mobile agents with rotational freedom, divided into two populations with…

Soft Condensed Matter · Physics 2025-11-20 Alessandro Scirè

Suspensions of swimming micro-organisms provide examples of coordinated active dynamics. That has stimulated the study of a phenomenological theory combining synchronization and polar order in active matter. Here, we consider another…

Soft Condensed Matter · Physics 2019-04-08 M. Leoni

A nematic liquid crystal confined to the surface of a sphere exhibits topological defects of total charge $+2$ due to the topological constraint. In equilibrium, the nematic field forms four $+1/2$ defects, located at the corners of a…

Soft Condensed Matter · Physics 2020-07-22 Yi-Heng Zhang , Markus Deserno , Zhan-Chun Tu

In biological systems, active agents such as actomyosin and cells move and interact on curved surfaces, exhibiting diverse phenomena. These observations have motivated studies of how curvature shapes their collective behavior. Here, using a…

Soft Condensed Matter · Physics 2025-12-19 Tatsuo Shibata

Active matter consists of units that generate mechanical work by consuming energy. Examples include living systems, such as assemblies of bacteria and biological tissues, biopolymers driven by molecular motors, and suspensions of synthetic…

Soft Condensed Matter · Physics 2021-02-04 Song Liu , Suraj Shankar , M. Cristina Marchetti , Yilin Wu

Remarkably, even under negligible inertia, the addition of microstructural agents can generate chaotic flow fields. Such behavior can arise in polymer solutions, leading to elastic turbulence, or from active, self-driven particles, which…

Fluid Dynamics · Physics 2026-01-15 Vedad Dzanic , Sumesh P. Thampi , Julia M. Yeomans