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Many microorganisms and artificial microswimmers use helical appendages in order to generate locomotion. Though often rotated so as to produce thrust, some species of bacteria such Spiroplasma, Rhodobacter sphaeroides and Spirochetes induce…

Biological Physics · Physics 2018-10-23 Lyndon Koens , Hang Zhang , Martin Moeller , Ahmed Mourran , Eric Lauga

Undulatory locomotion is a means of self-propulsion that relies on the generation and propagation of waves along a body. As a mode of locomotion it is primitive and relatively simple, yet can be remarkably robust. No wonder then, that it is…

Biological Physics · Physics 2009-08-20 Netta Cohen , Jordan H. Boyle

We develop a continuum theory of linear viscoelastic response in oriented monodomain nematic elastomers. The expression for dissipation function is analogous to the Leslie-Ericksen version of anisotropic nematic viscosity; we propose the…

Soft Condensed Matter · Physics 2009-11-07 E. M. Terentjev , M. Warner

Swimming micro-organisms such as flagellated bacteria and sperm cells have fascinating locomotion capabilities. Inspired by their natural motion, there is an ongoing effort to develop artificial robotic nano-swimmers for potential in-body…

Fluid Dynamics · Physics 2023-05-31 Jithu Paul , Yizhar Or , Oleg Gendelman

Many biological microswimmers can modulate their swimming gait to achieve directional control of motility, especially when performing steering towards specific directional cues. This can be achieved without the need for obvious…

Microswimmers are active particles of microscopic size that self-propel by setting the surrounding fluid into motion. According to the kind of far-field fluid flow that they induce, they are classified into pushers and pullers. Many studies…

Soft Condensed Matter · Physics 2019-08-01 Giorgio Pessot , Hartmut Löwen , Andreas M. Menzel

Both natural and artificial small-scale swimmers may often self-propel in environments subject to complex geometrical constraints. While most past theoretical work on low-Reynolds number locomotion addressed idealised geometrical…

Fluid Dynamics · Physics 2017-11-16 Alexander Chamolly , Takuji Ishikawa , Eric Lauga

Elongate animals and robots use undulatory body waves to locomote through diverse environments. Geometric mechanics provides a framework to model and optimize such systems in highly damped environments, connecting a prescribed shape change…

Robotics · Computer Science 2025-10-21 Jianfeng Lin , Tianyu Wang , Baxi Chong , Matthew Fernandez , Zhaochen Xu , Daniel I. Goldman

Both animal and plant tissue exhibit a nonlinear rheological phenomenon known as compression stiffening, or an increase in moduli with increasing uniaxial compressive strain. Does such a phenomenon exist in single cells, which are the…

We compute the temporal evolution of the pressure anisotropy and bulk pressure of a massive gas using second-order viscous hydrodynamics and anisotropic hydrodynamics. We then compare our results with an exact solution of the Boltzmann…

High Energy Physics - Phenomenology · Physics 2014-10-16 Gabriel S. Denicol , Wojciech Florkowski , Radoslaw Ryblewski , Michael Strickland

Ribbons are long narrow strips possessing three distinct material length scales (thickness, width, and length) which allow them to produce unique shapes unobtainable by wires or filaments. For example when a ribbon has half a twist and is…

Fluid Dynamics · Physics 2016-02-04 Lyndon Koens , Eric Lauga

A single flexible filament can be actuated to escape from the scallop theorem and generate net propulsion at low Reynolds number. In this work, we study the dynamics of a simple boundary-driven multi-filament swimmer, a two-arm clamshell…

Fluid Dynamics · Physics 2022-03-07 Shiyuan Hu , Jun Zhang , Michael J. Shelley

Actuating periodically an elastic filament in a viscous liquid generally breaks the constraints of Purcell's scallop theorem, resulting in the generation of a net propulsive force. This observation suggests a method to design simple…

Soft Condensed Matter · Physics 2009-09-29 Eric Lauga

This work comprises a detailed theoretical and computational study of the boundary value problem for transversely isotropic linear elastic bodies. General conditions for well-posedness are derived in terms of the material parameters. The…

Numerical Analysis · Mathematics 2018-11-01 Faraniaina Rasolofoson , Beverley Grieshaber , B. Daya Reddy

Incompressibility is established for three-dimensional and two-dimensional deformations of an anisotropic linearly elastic material, as conditions to be satisfied by the elastic compliances. These conditions make it straightforward to…

Soft Condensed Matter · Physics 2013-05-23 Michel Destrade , Paul A. Martin , Tom C. T. Ting

Using explicit-water molecular dynamics (MD) simulations of a generic pocket-ligand model we investigate how chemical and shape anisotropy of small ligands influences the affinities, kinetic rates and pathways for their association to…

Soft Condensed Matter · Physics 2020-09-18 R. Gregor Weiß , Richard Chudoba , Piotr Setny , Joachim Dzubiella

In this paper, we study the anisotropy classes of the fourth order elastic tensors of the relaxed micromorphic model, also introducing their second order counterpart by using a Voigt-type vector notation. In strong contrast with the usual…

Taylor's swimming sheet is a classical model of microscale propulsion and pumping. Many biological fluids and substances are fibrous, having a preferred direction in their microstructure; for example cervical mucus is formed of polymer…

Fluid Dynamics · Physics 2017-03-08 Gemma Cupples , Rosemary J. Dyson , David J. Smith

Dynamics at low Reynolds numbers experiences recent revival in the fields of biophysics and active matter. While in bulk isotropic fluids it is exhaustively studied, this is less so in anisotropic fluids and in confined situations. Here, we…

Soft Condensed Matter · Physics 2026-01-21 Abdallah Daddi-Moussa-Ider , Elsen Tjhung , Marc Pradas , Thomas Richter , Andreas M. Menzel

The random walk of a bonded monomer in a polymer melt is anisotropic due to local order and bond connectivity. We investigate both effects by molecular-dynamics simulations on melts of fully-flexible linear chains ranging from dimers…

Soft Condensed Matter · Physics 2016-05-06 S. Bernini , D. Leporini