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I formulate the theory of three dimensional "Malthusian flocks" -- i.e., coherently moving collections of self-propelled entities (such as living creatures) which are being "born" and "dying" during their motion -- whose constituents all…

Soft Condensed Matter · Physics 2024-11-14 John Toner

We study the hydrodynamic behavior of two-dimensional chiral dry Malthusian flocks; that is, chiral polar-ordered active matter with neither number nor momentum conservation. We show that, in the absence of fluctuations, such systems…

Soft Condensed Matter · Physics 2025-07-29 Leiming Chen , Chiu Fan Lee , John Toner

We propose a hydrodynamic description of the homogeneous ordered phase of polar flocks. Starting from symmetry principles, we construct the appropriate equation for the dynamics of the Goldstone mode associated with the broken rotational…

Statistical Mechanics · Physics 2024-03-07 Hugues Chaté , Alexandre Solon

I study "Malthusian Flocks": moving aggregates of self-propelled entities (e.g., organisms, cytoskeletal actin, microtubules in mitotic spindles) that reproduce and die. Long-ranged order (i.e., the existence of a non-zero average velocity…

Other Condensed Matter · Physics 2013-05-30 John Toner

We show that "Malthusian flocks" -- i.e., coherently moving collections of self-propelled entities (such as living creatures) which are being "born" and "dying" during their motion -- belong to a new universality class in spatial dimensions…

Soft Condensed Matter · Physics 2020-09-02 Leiming Chen , Chiu Fan Lee , John Toner

We show that "Malthusian flocks" -- i.e., coherently moving collections of self-propelled entities (such as living creatures) which are being "born" and "dying" during their motion -- belong to a new universality class in spatial dimensions…

Soft Condensed Matter · Physics 2020-09-02 Leiming Chen , Chiu Fan Lee , John Toner

We construct the hydrodynamic theory of coherent collective motion ("flocking") at a solid-liquid interface. The polar order parameter and concentration of a collection of "active" (self-propelled) particles at a planar interface between a…

Soft Condensed Matter · Physics 2022-01-05 Niladri Sarkar , Abhik Basu , John Toner

Although human beings have known about the phenomenon of "flocking"- that is, the coherent movement of large numbers of creatures (flocks of birds, schools of fish, herds of woolly mammoths, etc.)- since prehistoric times, it is only in the…

Soft Condensed Matter · Physics 2018-12-04 John Toner

I reanalyze the hydrodynamic theory of fluid, polar ordered flocks. I find new linear terms in the hydrodynamic equations which slightly modify the anisotropy, but not the scaling, of the damping of sound modes. I also find that the…

Other Condensed Matter · Physics 2015-06-04 John Toner

We study the moving phase of two-dimensional (2D) incompressible polar active fluids in the presence of both quenched and annealed disorder. We show that long-range polar order persists even in this defect-ridden two-dimensional system. We…

Soft Condensed Matter · Physics 2022-11-04 Leiming Chen , Chiu Fan Lee , Ananyo Maitra , John Toner

We study two-dimensional chiral dry Malthusian flocks; that is, chiral polar-ordered active matter with neither number nor momentum conservation. In the absence of fluctuations, these form a ``time cholesteric", in which the velocity…

Soft Condensed Matter · Physics 2025-07-09 Leiming Chen , Chiu Fan Lee , John Toner

We study a collection of polar self-propelled particles or polar flock on a two dimensional substrate with birth and death. Most of the previous studies of polar flock with birth and death have assumed the compressible flock, such that the…

Soft Condensed Matter · Physics 2022-12-16 Pratikshya Jena , Shradha Mishra

When interacting motile units self-organize into flocks, they realize one of the most robust ordered state found in nature. However, after twenty five years of intense research, the very mechanism controlling the ordering dynamics of both…

Soft Condensed Matter · Physics 2021-10-04 Amélie Chardac , Ludwig A. Hoffmann , Yoann Poupart , Luca Giomi , Denis Bartolo

We present the hydrodynamic theory of coherent collective motion ("flocking") at a solid-liquid interface, and many of its predictions for experiment. We find that such systems are stable, and have long-range orientational order, over a…

Soft Condensed Matter · Physics 2022-01-05 Niladri Sarkar , Abhik Basu , John Toner

We present a hydrodynamic model of flocking that generalizes the familiar Toner-Tu equations to incorporate turning inertia of well-polarized flocks. The continuum equations controlled by only two dimensionless parameters, orientational…

Soft Condensed Matter · Physics 2016-01-20 Xingbo Yang , M. Cristina Marchetti

The hydrodynamic equations of dry active polar fluids (i.e., moving flocks without momentum conservation) are shown to imply giant number fluctuations. Specifically, the rms fluctuations $\sqrt {<(\delta N)^2>}$ of the number $N$ of active…

Statistical Mechanics · Physics 2019-05-22 John Toner

We show that incompressible polar active fluids can exhibit an ordered, coherently moving phase even in the presence of quenched disorder in two dimensions. Unlike such active fluids with annealed (i.e., time-dependent) disorder only, which…

Soft Condensed Matter · Physics 2022-11-04 Leiming Chen , Chiu Fan Lee , Ananyo Maitra , John Toner

We investigate the effect of cooperative interactions in an ensemble of microorganisms, modelled as self-propelled disk-like and rod-like particles, in a three-dimensional turbulent flow to show flocking as an emergent phenomenon. Building…

Soft Condensed Matter · Physics 2020-05-20 Anupam Gupta , Amal Roy , Arnab Saha , Samriddhi Sankar Ray

We present a hydrodynamic theory of incompressible polar active fluids with quenched disorder. This theory shows that such fluids can overcome the disruption caused by the quenched disorder and move coherently, in the sense of having a…

Soft Condensed Matter · Physics 2022-11-04 Leiming Chen , Chiu Fan Lee , Ananyo Maitra , John Toner

We present a comprehensive theory of the dynamics and fluctuations of a two-dimensional suspension of polar active particles in an incompressible fluid confined to a substrate. We show that, depending on the sign of a single parameter, a…

Soft Condensed Matter · Physics 2020-01-22 Ananyo Maitra , Pragya Srivastava , M. Cristina Marchetti , Sriram Ramaswamy , Martin Lenz
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