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Spatial distribution of the human population is distinctly heterogeneous, e.g. showing significant difference in the population density between urban and rural areas. In the historical perspective, i.e. on the timescale of centuries, the…

Adaptation and Self-Organizing Systems · Physics 2022-08-30 Anna Zincenko , Sergei Petrovskii , Vitaly Volpert

We study the mechanisms of pattern formation for vegetation dynamics in water-limited regions. Our analysis is based on a set of two partial differential equations (PDEs) of reaction-diffusion type for the biomass and water and one ordinary…

Dynamical Systems · Mathematics 2023-03-24 Konstantinos Spiliotis , Lucia Russo , Francesco Giannino , Constantinos Siettos

Several mechanisms have been proposed to explain the spontaneous generation of self-organized patterns, hypothesised to play a role in the formation of many of the magnificent patterns observed in Nature. In several cases of interest, the…

Pattern Formation and Solitons · Physics 2025-10-22 Riccardo Muolo , Malbor Asllani , Duccio Fanelli , Philip K. Maini , Timoteo Carletti

Among living organisms, there are species that change their patterns on their body surface during their growth process and those that maintain their patterns. Theoretically, it has been shown that large-scale species do not form distinct…

Biological Physics · Physics 2025-08-27 Shin Nishihara , Toru Ohira

We performed an extensive numerical study of a two-dimensional reaction-diffusion system of the activator-inhibitor type in which domain patterns can form. We showed that both multidomain and labyrinthine patterns may form spontaneously as…

patt-sol · Physics 2016-09-08 C. B. Muratov , V. V. Osipov

Spatial self-organization emerges in distributed systems exhibiting local interactions when nonlinearities and the appropriate propagation of signals are at work. These kinds of phenomena can be modeled with different frameworks, typically…

Cell Behavior · Quantitative Biology 2016-11-23 Adriano Bonforti , Salva Duran-Nebreda , Raul Montañez , Ricard Solé

Turing patterns are a central paradigm for describing spatial patterns in nature. The corresponding theory of reaction-diffusion dynamics combines ideal diffusion with nonlinear reactions, resulting in patterns when species diffuse at…

Biological Physics · Physics 2026-01-28 Cathelijne ter Burg , David Zwicker

The rheological properties of biological tissues are core to processes such as cancer metastasis, wound healing and embryo development. The emergence of tissue and organ structures during morphogenesis requires the precise formation of…

Soft Condensed Matter · Physics 2026-03-10 Aidan J. Nicholas , Suzanne M. Fielding

Some quantities in the reaction-diffusion models from cellular biology or ecology depend on the spatial average of density functions instead of local density functions. We show that such nonlocal spatial average can induce instability of…

Analysis of PDEs · Mathematics 2020-02-03 Qingyan Shi , Junping Shi , Yongli Song

The aim of this paper is to contribute to the understanding of the pattern formation phenomenon in reaction-diffusion equations coupled with ordinary differential equations. Such systems of equations arise, for example, from modeling of…

Analysis of PDEs · Mathematics 2016-07-15 Anna Marciniak-Czochra , Grzegorz Karch , Kanako Suzuki

Several theoretical models predict that spatial patterning increases ecosystem resilience. However, these predictions rely on simplifying assumptions, such as assuming isotropic and infinitely large ecosystems, and empirical evidence…

Populations and Evolution · Quantitative Biology 2026-04-06 David Pinto-Ramos , Ricardo Martinez-Garcia

In this paper the Turing pattern formation mechanism of a two component reaction-diffusion system modeling the Schnakenberg chemical reaction coupled to linear cross-diffusion terms is studied. The linear cross-diffusion terms favors the…

Pattern Formation and Solitons · Physics 2017-05-08 G. Gambino , S. Lupo , M. Sammartino

A feature common to many models of vegetation pattern formation in semi-arid ecosystems is a sequence of qualitatively different patterned states, "gaps -> labyrinth -> spots", that occurs as a parameter representing precipitation…

Pattern Formation and Solitons · Physics 2014-10-10 Karna Gowda , Hermann Riecke , Mary Silber

Spatial patterns are widely observed in numerous nonequilibrium natural systems, often undergoing complex transitions and bifurcations, thereby exhibiting significant importance in many physical and biological systems such as embryonic…

Biological Physics · Physics 2024-12-17 Jie Su , Wei Wu , Denis Patterson , Simon Asher Levin , Jin Wang

We study the movement of the living organism in a band form towards the presence of chemical substrates based on a system of partial differential evolution equations. We incorporate Einstein's method of Brownian motion to deduce the…

Analysis of PDEs · Mathematics 2023-10-10 Rahnuma Islam , Akif Ibragimov

Accurate and robust spatial orders are ubiquitous in living systems. In 1952, Alan Turing proposed an elegant mechanism for pattern formation based on spontaneous breaking of the spatial translational symmetry in the underlying…

Statistical Mechanics · Physics 2022-06-07 Dongliang Zhang , Chenghao Zhang , Qi Ouyang , Yuhai Tu

Self-organization, the ability of a system of microscopically interacting entities to shape macroscopically ordered structures, is ubiquitous in Nature. Spatio-temporal patterns are abundantly observed in a large plethora of applications,…

Pattern Formation and Solitons · Physics 2019-06-17 Malbor Asllani , Timoteo Carletti , Duccio Fanelli , Philip K. Maini

General conditions are established under which reaction-cross-diffusion systems can undergo spatiotemporal pattern-forming instabilities. Recent work has focused on designing systems theoretically and experimentally to exhibit patterns with…

Pattern Formation and Solitons · Physics 2025-08-26 Edgardo Villar-Sepúlveda , Alan R. Champneys , Andrew L. Krause

Biological systems excel at building spatial structures on scales ranging from nanometers to kilometers and exhibit temporal patterning from milliseconds to years. One approach that nature has taken to accomplish this relies on the…

Cell Behavior · Quantitative Biology 2009-11-10 Herbert Levine , Eshel Ben-Jacob

We identify a new type of pattern formation in spatially distributed active systems. We simulate one-dimensional two-component systems with predator-prey local interaction and pursuit-evasion taxis between the components. In a sufficiently…

Pattern Formation and Solitons · Physics 2013-05-29 V. N. Biktashev , M. A. Tsyganov