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Researchers have long debated which spatial arrangements and swimming synchronizations are beneficial for the hydrodynamic performance of fish in schools. In our previous work (Seo and Mittal, Bioinsp. Biomim., Vol. 17, 066020, 2022), we…

Fluid Dynamics · Physics 2025-07-11 Ji Zhou , Jung-Hee Seo , Rajat Mittal

Fish schooling is often modeled with self-propelled particles subject to phenomenological behavioral rules. Although fish are known to sense and exploit flow features, these models usually neglect hydrodynamics. Here, we propose a novel…

Biological Physics · Physics 2018-05-16 Audrey Filella , François Nadal , Clément Sire , Eva Kanso , Christophe Eloy

The complex formations exhibited by schooling fish have long been the object of fascination for biologists and physicists. However, the physical and sensory mechanisms leading to organized collective behavior remain elusive. On the physical…

Fluid Dynamics · Physics 2025-07-09 Anand U. Oza , Eva Kanso , Michael J. Shelley

There is increasing evidence that fish gain energetic benefits when they swim in a school. The most recent indications of such benefits are a lower tail (or fin) beat at the back of a school and reduced oxygen consumption in schooling fish…

Quantitative Methods · Quantitative Biology 2013-07-30 C. K. Hemelrijk , D. A. P. Reid , H. Hildenbrandt , J. T. Padding

Schooling, an archetype of collective behavior, emerges from the interactions of fish responding to visual and other informative cues mediated by their aqueous environment. In this context, a fundamental and largely unexplored question…

Living fish may suddenly encounter upstream obstacles, join the queue of the fish schooling, or detect upstream flow in advance, resulting in interactions with environmental vortices that can be abrupt or develop gradually from an initial…

Fluid Dynamics · Physics 2024-09-27 Peng Han , Dong Zhang , Jun-Duo Zhang , Wei-Xi Huang

Fish perform various propulsive maneuvers while swimming by generating traveling waves along their bodies and producing thrust through tail strokes. Anguilliform swimmers spread motion along the body, while carangiform swimmers' motion is…

Fluid Dynamics · Physics 2025-12-04 Zahra Maleksabet , Maham Kamran , Ali Tarokh , Muhammad Saif Ullah Khalid

Three-dimensional experiments are presented on a school of three pitching hydrofoils. Two side-by-side leader foils maintain the same relative positions while the location of a third follower foil is varied. Force and flow measurements…

Fluid Dynamics · Physics 2026-04-23 Pedro C. Ormonde , Yuanhang Zhu , Daniel Quinn , Keith W Moored

Fish schools are ubiquitous in marine life. Although flow interactions are thought to be beneficial for schooling, their exact effects on the speed, energetics, and stability of the group remain elusive. Recent experiments suggest that flow…

Fluid Dynamics · Physics 2021-08-04 Sina Heydari , Eva Kanso

Fish in schooling formations navigate complex flow-fields replete with mechanical energy in the vortex wakes of their companions. Their schooling behaviour has been associated with evolutionary advantages including collective energy…

Fluid Dynamics · Physics 2022-06-08 Siddhartha Verma , Guido Novati , Petros Koumoutsakos

Many species of fish, as well as biorobotic underwater vehicles, employ body caudal fin propulsion, in which a wave-like body motion culminates in high-amplitude caudal fin oscillations to generate thrust. This study uses high fidelity…

Fluid Dynamics · Physics 2026-03-03 Jung Hee Seo , Ji Zhou , Rajat Mittal

We present new constrained and free-swimming experiments and simulations of a pair of pitching hydrofoils interacting in a minimal school. The hydrofoils have an out-of-phase synchronization and they are varied through in-line, staggered,…

Fluid Dynamics · Physics 2023-08-01 Pedro Costa Ormonde , Melike Kurt , Amin Mivehchi , Keith W. Moored

It has been a long-standing problem how schooling fish optimize their motion by exploiting the vortices shed by the others. A recent experimental study showed that a pair of fish reduce energy consumption by matching the phases of their…

Biological Physics · Physics 2023-12-05 Susumu Ito , Nariya Uchida

Fish schools and bird flocks exhibit complex collective dynamics whose self-organization principles are largely unknown. The influence of hydrodynamics on such collectives has been relatively unexplored theoretically, in part due to the…

Soft Condensed Matter · Physics 2019-11-06 Anand U. Oza , Leif Ristroph , Michael J. Shelley

Inspired by the wake-surfing nature of animals, this study aims to understand the aerodynamic force variation on a wing surfing in an unsteady 2-D wake. Wind tunnel experiments were conducted using Particle Image Velocimetry (PIV) and force…

Fluid Dynamics · Physics 2025-12-11 Siyang Hao , Kenneth Breuer

Fish often travel in highly organized schools. One of the most quoted functions of these configurations is energy savings. Here, we verified the hypothesis and explored the mechanism through series of experiments on "schooling" robotic…

Biological Physics · Physics 2015-08-14 Liang Li , Lichao Jia , Guangming Xie

Fish typically swim by periodic bending of their bodies. Bending seems to follow a universal rule; it occurs at about one-third from the posterior end of the fish body with a maximum bending angle of about $30^o$. However, the hydrodynamic…

Fluid Dynamics · Physics 2021-12-22 Haotian Hang , Sina Heydari , John H. Costello , Eva Kanso

One of the most highly debated questions in the field of animal swarming and social behaviour, is the collective random patterns and chaotic behaviour formed by some animal species, in particular if there is a danger. Is such a behaviour…

Populations and Evolution · Quantitative Biology 2017-01-04 Usama Kadri , Franz Brümmer , Anan Kadri

Wake sensing for bioinspired robotic swimmers has been the focus of much investigation owing to its relevance to locomotion control, especially in the context of schooling and target following. Many successful wake sensing strategies have…

Fluid Dynamics · Physics 2017-11-30 Mengying Wang , Maziar S. Hemati

Inspired by dense contractile tissues, where cells are subject to periodic deformation, we formulate and study a generic hydrodynamic theory of pulsating active liquids. Combining mechanical and phenomenological arguments, we postulate that…

Soft Condensed Matter · Physics 2025-09-25 Tirthankar Banerjee , Thibault Desaleux , Jonas Ranft , Étienne Fodor
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