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Phase separation driven by nonequilibrium fluctuations is a hallmark of both living and synthetic active matter. Unlike equilibrium systems, where ordered states arise from the minimization of free energy, active systems are fueled by a…

Active solids consist of elastically coupled out-of-equilibrium units performing work. They are central to autonomous processes, such as locomotion, self-oscillations and rectification, in biological systems,designer materials and robotics.…

Soft Condensed Matter · Physics 2025-01-17 P. Baconnier , D. Shohat , C. Hernandèz , C. Coulais , V. Démery , G. Düring , O. Dauchot

Flocking phase transitions found in models of polar active matter are paradigmatic examples of active phase transitions in soft matter. An interesting specialization of flocking models concerns a ``topological'' vs ``metric'' choice by…

Soft Condensed Matter · Physics 2024-09-10 Charles R. Packard , Daniel M. Sussman

Collective cell migration is crucial in many biological processes such as wound healing, tissue morphogenesis, and tumor progression. The leading front of a collective migrating epithelial cell layer often destabilizes into multicellular…

Biological Physics · Physics 2020-05-07 Yanjun Yang , Herbert Levine

The dynamics of dry active matter have implications for a diverse collection of biological phenomena spanning a range of length and time scales, such as animal flocking, cell tissue dynamics, and swarming of inserts and bacteria. Uniting…

Soft Condensed Matter · Physics 2021-01-07 David Nesbitt , Gunnar Pruessner , Chiu Fan Lee

Coherent vortical motion has been reported in a wide variety of populations including living organisms (bacteria, fishes, human crowds) and synthetic active matter (shaken grains, mixtures of biopolymers), yet a unified description of the…

With increasing emphasis on the study of active solids, the features of these classes of nonequilibrium systems and materials beyond their mere existence shift into focus. One concept of active solids addresses them as active,…

Soft Condensed Matter · Physics 2025-02-10 Yutaka Kinoshita , Nariya Uchida , Andreas M. Menzel

Many types of motile cells perform durotaxis, namely, directed migration following gradients of substrate stiffness. Recent experiments have revealed that cell monolayers can migrate toward stiffer regions even when individual cells do not…

Biological Physics · Physics 2022-01-26 Irina Pi-Jaumà , Ricard Alert , Jaume Casademunt

Two hallmarks of non-equilibrium systems, from active colloids to animal herds, are agents motility and nonreciprocal interactions. Their interplay creates feedback loops leading to complex spatiotemporal dynamics crucial to understand and…

Statistical Mechanics · Physics 2025-09-17 David Martin , Daniel Seara , Yael Avni , Michel Fruchart , Vincenzo Vitelli

We show that the flocking transition in the Vicsek model is best understood as a liquid-gas transition, rather than an order-disorder one. The full phase separation observed in flocking models with Z2 rotational symmetry is, however,…

Statistical Mechanics · Physics 2015-04-03 Alexandre P. Solon , Hugues Chaté , Julien Tailleur

We formulate a hydrodynamic theory of confluent epithelia: i.e. monolayers of epithelial cells adhering to each other without gaps. Taking advantage of recent progresses toward establishing a general hydrodynamic theory of p-atic liquid…

Soft Condensed Matter · Physics 2024-01-11 Josep-Maria Armengol-Collado , Livio Nicola Carenza , Luca Giomi

We present a comprehensive computational study of the collective behavior emerging from the competition between self-propulsion, excluded volume interactions and velocity-alignment in a two-dimensionnal model of active particles. We…

Soft Condensed Matter · Physics 2018-01-08 Aitor Martín-Gómez , Demian Levis , Albert Díaz-Guilera , Ignacio Pagonabarraga

Flocking behavior is observed in biological systems from the cellular to super-organismal length scales, and the mechanisms and purposes of this behavior are objects of intense interest. In this paper, we study the collective dynamics of…

We present a hydrodynamic model of spreading epithelial monolayers as polar viscous fluids, with active contractility and traction on the substrate. The combination of both active forces generate an instability that leads to nonlinear…

Soft Condensed Matter · Physics 2017-05-02 C. Blanch-Mercader , J. Casademunt

We study the spatio-temporal dynamics of a model of polar active fluid in two dimensions. The system exhibits a transition from an isotropic to a polarized state as a function of density. The uniform polarized state is, however, unstable…

Soft Condensed Matter · Physics 2011-12-08 Luca Giomi , M. Cristina Marchetti

Active solids such as cell collectives, colloidal clusters, and active metamaterials exhibit diverse collective phenomena, ranging from rigid body motion to shape-changing mechanisms. The nonlinear dynamics of such active materials remains…

Soft Condensed Matter · Physics 2024-06-21 Claudio Hernández-López , Paul Baconnier , Corentin Coulais , Olivier Dauchot , Gustavo Düring

All types of blood cells are formed by differentiation from a small self-maintaining population of pluri-potential stem cells in the bone marrow. Despite abundant information on the molecular aspects of division, differentiation, commitment…

Biological Physics · Physics 2007-05-23 Carlos A. Perazzo , Elmer A. Fernandez , Dante R. Chialvo , Peter Willshaw

Cells not only can be motile by crawling but are also capable of non-motility active motions like periodic contraction or pulsation. In this work, based on a Voronoi cell model, we show how this non-motility activity affects the structure,…

Soft Condensed Matter · Physics 2025-02-11 Zhu-Qin Li , Qun-Li Lei , Yu-qiang Ma

Confluent cell monolayers and epithelia tissues show remarkable patterns and correlations in structural arrangements and actively-driven collective flows. We simulate these properties using multiphase field models. The models are based on…

Soft Condensed Matter · Physics 2021-12-08 Dennis Wenzel , Axel Voigt

In an inspiring recent study, Xu et al. [Nat. Phys. 19, 46 (2023)] observed for a living active biofilm under circular confinement two emergent dynamic modes of collective motion in the film. One corresponds to global rotational motion of…

Soft Condensed Matter · Physics 2025-02-11 Henning Reinken , Andreas M. Menzel