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We discuss a simple model of particles hopping in one dimension with attractive interactions. Taking a hydrodynamic limit in which the interaction strength increases with the system size, we observe the formation of multiple clusters of…

Statistical Mechanics · Physics 2017-05-05 Matthew Burman , Daniel Carpenter , Robert L. Jack

We show that a two-dimensional system of flocking microswimmers interacting hydrodynamically can be expressed using a Hamiltonian formalism. The Hamiltonian depends strictly on the angles between the particles and their swimming…

Soft Condensed Matter · Physics 2023-05-23 Yuval Shoham , Naomi Oppenheimer

Granular materials are inherently out-of-equilibrium systems due to energy dissipation through inelastic collisions and friction. When driven by mechanical agitation such as vibration, they exhibit rich collective behaviors including…

Soft Condensed Matter · Physics 2026-05-01 Kai Kono , Hiroyuki Ebata , Shio Inagaki

Swimming cells and microorganisms are as diverse in their collective dynamics as they are in their individual shapes and propulsion mechanisms. Even for sperm cells, which have a stereotyped shape consisting of a cell body connected to a…

Fluid Dynamics · Physics 2018-06-25 S. F. Schoeller , E. E. Keaveny

Suspensions of swimming micro-organisms are known to undergo intricate collective dynamics as a result of hydrodynamic and collision interactions. Micro-swimmers, such as bacteria and micro-algae, naturally live and have evolved in complex…

Fluid Dynamics · Physics 2025-10-31 Yasser Almoteri , Enkeleida Lushi

Recent experiments on monolayers of spindle-like cells plated on adhesive stripe-shaped domains have provided a convincing demonstration that certain types of collective phenomena in epithelia are well described by active nematic…

Soft Condensed Matter · Physics 2020-01-22 Ludwig A. Hoffmann , Koen Schakenraad , Roeland M. H. Merks , Luca Giomi

Active colloids are microscopic particles, which self-propel through viscous fluids by converting energy extracted from their environment into directed motion. We first explain how articial microswimmers move forward by generating…

Soft Condensed Matter · Physics 2016-05-12 Andreas Zöttl , Holger Stark

Systems of self-propelled particles are known for their tendency to aggregate and to display swarm behavior. We investigate two model systems, self-propelled rods interacting via volume exclusion, and sinusoidally-beating flagella embedded…

Biological Physics · Physics 2010-09-28 Yingzi Yang , Vincent Marceau , Gerhard Gompper

Self-propelled particles move along circles rather than along a straight line when their driving force does not coincide with their propagation direction. Examples include confined bacteria and spermatozoa, catalytically driven nanorods,…

Soft Condensed Matter · Physics 2008-08-18 Sven van Teeffelen , Hartmut Löwen

Swimming microorganisms can influence the diffusion of passive particles. The effect of this swimmer-particle interaction depends on different properties, such as the hydrodynamic field of the swimmer and the relative sizes of…

Soft Condensed Matter · Physics 2020-05-08 Florian von Rüling , Francine Kolley , Alexey Eremin

Living organisms exhibit consistent homochirality. It is argued that the specific, binary choice made is not an accident but is a consequence of parity violation in the weak interaction expressed by cosmic irradiation. The secondary muons…

Biological Physics · Physics 2019-11-07 Noemie Globus , Roger D. Blandford

For natural microswimmers, the interplay of swimming activity and external flow can promote robust motion, e.g. propulsion against ("upstream rheotaxis") or perpendicular to the direction of flow. These effects are generally attributed to…

Soft Condensed Matter · Physics 2018-01-30 Jaideep Katuri , William E. Uspal , Juliane Simmchen , Albert Miguel-López , Samuel Sánchez

Cluster growth in a coagulating system of active particles (such as microswimmers in a solvent) is studied by theory and simulation. In contrast to passive systems, the net velocity of a cluster can have various scalings dependent on the…

Soft Condensed Matter · Physics 2014-03-21 P. Cremer , H. Löwen

Many active biological particles, such as swimming microorganisms or motor-proteins, do work on their environment by going though a periodic sequence of shapes. Interactions between particles can lead to the phase-synchronization of their…

Soft Condensed Matter · Physics 2022-07-26 Brato Chakrabarti , Michael J. Shelley , Sebastian Fürthauer

Bacteria exist in a free-swimming state or in a sessile biofilm state. The transition from free-swimming to sessile mode is characterized by changes in gene expression which alter, among others,the mechanics of flagellar motility. In this…

Fluid Dynamics · Physics 2014-05-07 Alan Cheng Hou Tsang , Eva Kanso

We numerically investigate the motion of active artificial microswimmers diffusing in a fuel concentration gradient. We observe that, in the steady state, their probability density accumulates in the low-concentration regions, whereas a…

Soft Condensed Matter · Physics 2025-08-18 Pulak K. Ghosh , Yunyun Li , Fabio Marchesoni , Franco Nori

The collective behavior of active semiflexible filaments is studied with a model of tangentially driven self-propelled worm-like chains. The combination of excluded-volume interactions and self-propulsion leads to several distinct dynamic…

Biological Physics · Physics 2024-06-03 Ozer Duman , Rolf E. Isele-Holder , Jens Elgeti , Gerhard Gompper

We investigate the collective transport properties of microscopic magnetic rollers that propel close to a surface due to a circularly polarized, rotating magnetic field. The applied field exerts a torque to the particles, which induces a…

Soft Condensed Matter · Physics 2022-01-03 Gaspard Junot , Andrejs Cebers , Pietro Tierno

We explore the interplay between tunneling process and chiral interactions in the discrimination of chiral states for an ensemble of molecules in a biological environment. Each molecule is described by an asymmetric double-well potential…

Biomolecules · Quantitative Biology 2017-04-28 Arash Tirandaz , Farhad Taher Ghahramani , Afshin Shafiee

We report computer simulation results on the cluster formation of dipolar colloidal particles driven by a rotating external field in a quasi-two-dimensional setup. We focus on the interplay between permanent dipolar and hydrodynamic…

Soft Condensed Matter · Physics 2015-06-12 Sebastian Jäger , Holger Stark , Sabine H. L. Klapp