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Living microorganisms are capable of a tactic response to external stimuli by swimming towards or away from the stimulus source; they do so by adapting their tactic signal transduction pathways to the environment. Their self-motility thus…

Soft Condensed Matter · Physics 2016-10-12 Alexander Geiseler , Peter Hänggi , Fabio Marchesoni , Colm Mulhern , Sergey Savel'ev

We study numerically the hydrodynamics of a self-propelled particle system, consisting of spherical squirmers sedimented on a flat surface. We observe the emergence of dynamic structures, due to the interplay of particle-particle and…

Soft Condensed Matter · Physics 2019-12-20 Zaiyi Shen , Juho S. Lintuvuori

Collective states of inanimate particles self-assemble through physical interactions and thermal motion. Despite some phenomenological resemblance, including signatures of criticality, the autonomous dynamics that binds motile agents into…

Soft Condensed Matter · Physics 2023-02-01 Xiangzun Wang , Pin-Chuan Chen , Klaus Kroy , Viktor Holubec , Frank Cichos

Population heterogeneity is ubiquitous among active living systems, but little is known about its role in determining their spatial organization and large-scale dynamics. Combining evidence from synthetic active fluids assembled from…

Soft Condensed Matter · Physics 2023-11-27 Samadarshi Maity , Alexandre Morin

A key goal in developing molecular microrobots that mimic real-world animal dynamic behavior is to understand better the self-continuous progressive motion resulting from collective molecular transformation. This study reports, for the…

Soft Condensed Matter · Physics 2025-05-09 Kazuma Obara , Yoshiyuki Kageyama , Sadamu Takeda

Active matter exhibits various forms of non-equilibrium states in the absence of external forcing, including macroscopic steady-state currents. Such states are often too complex to be modelled from first principles and our understanding of…

Soft Condensed Matter · Physics 2020-09-17 Viktor Škultéty , Cesare Nardini , Joakim Stenhammar , Davide Marenduzzo , Alexander Morozov

Fission and fusion processes of particles clusters occur in many areas of physics and chemistry from subnuclear to astronomic length scales. Here we study fission and fusion of magnetic microswimmer clusters as governed by their…

Soft Condensed Matter · Physics 2016-12-21 Francisca Guzmán-Lastra , Andreas Kaiser , Hartmut Löwen

Active colloids self-organise to a variety of collective states, ranging from highly motile 'molecules' to complex 3D structures. Using large-scale simulations, we show that hydrodynamic interactions, together with a gravity-like aligning…

Soft Condensed Matter · Physics 2020-03-31 Zaiyi Shen , Alois Würger , Juho S. Lintuvuori

Biological microswimmers such as bacteria and sperm cells often encounter complex biological fluid environments. Here we use the well-known squirmer microswimmer model to show the importance of the local fluid microstructure and…

Soft Condensed Matter · Physics 2023-07-19 Andreas Zöttl

We investigate density fluctuations in three-dimensional chiral active fluids by using a simple model of helical self-propelled particles. Helical motion is generated by a constant angular velocity (or chiral torque) acting on the…

Soft Condensed Matter · Physics 2025-10-29 Yuta Kuroda , Takeshi Kawasaki , Kunimasa Miyazaki

Lipid rafts are hypothesized to facilitate protein interaction, tension regulation, and trafficking in biological membranes, but the mechanisms responsible for their formation and maintenance are not clear. Insights into many other…

Soft Condensed Matter · Physics 2017-04-12 Louis Kang , T. C. Lubensky

Self-propelled micron-size particles suspended in a fluid, like bacteria or synthetic microswimmers, are strongly non-equilibrium systems where particle motility breaks the microscopic detailed balance, often resulting in large-scale…

Soft Condensed Matter · Physics 2024-03-15 Dóra Bárdfalvy , Viktor Škultéty , Cesare Nardini , Alexander Morozov , Joakim Stenhammar

Biological organisms often have elongated, flexible structures with some degree of chirality in their bodies or movements. In nature, these organisms frequently take advantage of self-encapsulation mechanisms that create folded…

Soft Condensed Matter · Physics 2024-10-04 Lorenzo Caprini , Iman Abdoli , Umberto Marini Bettolo Marconi , Hartmut Löwen

Self-propelled particles can spontaneously form dense phases from a dilute suspension in a process referred to as motility-induced phase separation. The properties of the out-of-equilibrium structures that are formed are governed by the…

Soft Condensed Matter · Physics 2016-09-08 Sven Erik Ilse , Christian Holm , Joost de Graaf

Collective behaviour in living systems is observed across many scales, from bacteria to insects, to fish shoals. Zebrafish have emerged as a model system amenable to laboratory study. Here we report a three-dimensional study of the…

Statistical Mechanics · Physics 2024-05-16 Yushi Yang , Francesco Turci , Erika Kague , Chrissy L. Hammond , John Russo , C. Patrick Royall

Collective behavior of self-propelled particles is observed on a microscale for swimmers such as sperm and bacteria as well as for protein filaments in motility assays. The properties of such systems depend both on their dimensionality and…

Soft Condensed Matter · Physics 2014-01-07 Masoud Abkenar , Kristian Marx , Thorsten Auth , Gerhard Gompper

Three-dimensional simulations with fully resolved hydrodynamics are performed to study the dynamics of a single squirmer under gravity, in order to clarify its motion in the vicinity of a flat plate. Different dynamics emerge for different…

Fluid Dynamics · Physics 2020-05-27 Federico Fadda , John Jairo Molina , Ryoichi Yamamoto

We present a hydrodynamic study of a monolayer of squirmer model microswimmers confined to a boundary by strong gravity using the simulation method of multi-particle collision dynamics. The squirmers interact with each other via their…

Soft Condensed Matter · Physics 2019-07-23 Jan-Timm Kuhr , Felix Rühle , Holger Stark

In this paper, we discuss microscopic models for chiral active particles, i.e., rotating active units that exhibit circular or spinning motion. While non-chiral active particles are typically governed by self-propulsion and conservative…

Soft Condensed Matter · Physics 2025-10-20 Lorenzo Caprini , Alessandro Petrini , Umberto Marini Bettolo Marconi

We present a comprehensive computational study of the collective behavior emerging from the competition between self-propulsion, excluded volume interactions and velocity-alignment in a two-dimensionnal model of active particles. We…

Soft Condensed Matter · Physics 2018-01-08 Aitor Martín-Gómez , Demian Levis , Albert Díaz-Guilera , Ignacio Pagonabarraga