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The bending of cilia and flagella is driven by forces generated by dynein motor proteins. These forces slide adjacent microtubule doublets within the axoneme, the motile cytoskeletal structure. To create regular, oscilla- tory beating…

Biological Physics · Physics 2015-06-19 V. Mukundan , P. Sartori , V. F. Geyer , F. Julicher , J. Howard

Microscale fluid flows generated by ensembles of beating eukaryotic flagella are crucial to fundamental processes such as development, motility and sensing. Despite significant experimental and theoretical progress, the underlying physical…

Soft Condensed Matter · Physics 2014-03-11 Douglas R. Brumley , Kirsty Y. Wan , Marco Polin , Raymond E. Goldstein

Micro-organisms usually can swim in their liquid environment by flagellar or ciliary beating. In this numerical work, we analyze the influence of flagellar beating on the orbits of a swimming cell in a shear flow. We also calculate the…

Soft Condensed Matter · Physics 2017-11-21 Levan Jibuti , Walter Zimmermann , Salima Rafaï , Philippe Peyla

Three-dimensional excitable systems can selforganize vortex patterns that rotate around one-dimensional phase singularities called filaments. In experiments with the Belousov-Zhabotinsky reaction and numerical simulations, we pin these…

Pattern Formation and Solitons · Physics 2015-04-08 Hua Ke , Zhihui Zhang , Oliver Steinbock

The beating flagella of the green alga Chlamydomonas reinhardtii play a prominent role in cellular mechanics, enabling cells to both displace and sense surrounding fluid. Specifically, flagellum-induced fluid transport enables microalgae to…

Fluid Dynamics · Physics 2026-03-10 Md Iftekhar Yousuf Emon , Gregorius R. Pradipta , Xiang Cheng , Xin Yong

We investigate a flexible polymer chain made up of chiral active Brownian particles in two dimensions using computer simulations. In the presence of chiral active Brownian forces, the radius of gyration of the chain reduces significantly.…

Soft Condensed Matter · Physics 2025-05-05 Shalabh K. Anand

A particle moving through a worm-like micellar fluid (WLM) shows instability and large fluctuations beyond a threshold \textcolor{black}{velocity/force (depending on the control parameter used)}. Despite many detailed studies, a direct…

Soft Condensed Matter · Physics 2025-12-15 Abhishek Ghadai , Pradip Kumar Bera , Sayantan Majumdar

Using simulation and theory we study the dynamics of a colloidal suspension in two dimensions subject to a time-delayed repulsive feedback that depends on the positions of the colloidal particles. The colloidal particles experience an…

Soft Condensed Matter · Physics 2019-09-06 Sonja Tarama , Stefan U. Egelhaaf , Hartmut Löwen

Flagella allow eukaryotic cells to move and pump fluid. We present the first three-dimensional, time-resolved imaging of the flagellar waveform of Chlamydomonas reinhardtii, a model alga found in fresh water. During the power stroke, we…

Biological Physics · Physics 2022-10-18 Laurence G. Wilson , Martin A. Bees

Collections of simple, self-propelled colloidal particles exhibit complex, emergent dynamical behavior, with promising applications in microrobotics. When confined within a deformable vesicle, self-propelled rods cluster and align,…

Soft Condensed Matter · Physics 2025-10-28 Sophie Y. Lee , Philipp W. A. Schönhöfer , Sharon C. Glotzer

In this paper, we present a theory to efficiently deal with mechanical properties of heterogeneous polymer chain in free space and the central problem is to evaluate the diffusion equation and orientation-orientation correlation function,…

Biological Physics · Physics 2019-11-05 Ming Li , Jizeng Wang

Three-dimensional excitable systems can create nonlinear scroll waves that rotate around one-dimensional phase singularities. Recent theoretical work predicts that these filaments drift along step-like height variations. Here we test this…

Pattern Formation and Solitons · Physics 2015-03-19 Hua Ke , Zhihui Zhang , Oliver Steinbock

Micro-scale cilia play a vital role in mucociliary clearance (MCC) in the human respiratory airways. In this numerical study, we examine fluid transport driven by the active beating of a single filament immersed in a three-dimensional…

Fluid Dynamics · Physics 2026-02-02 Qian Mao , Umberto d'Ortona , Julien Favier

Active polymers play a central role in many biological systems, from bacterial flagella to cellular cytoskeletons. Minimal models of semiflexible active filaments have been used to study a variety of interesting phenomena in active systems,…

Soft Condensed Matter · Physics 2020-02-19 Matthew S. E. Peterson , Michael F. Hagan , Aparna Baskaran

Mesh-like structures, such as mucus gel or cytoskeleton networks, are ubiquitous in biological systems. These intricate structures are composed of cross-linked, semi-flexible bio-filaments, crucial to numerous biological processes. In many…

Soft Condensed Matter · Physics 2024-01-10 Yeongjin Kim , Won Kyu Kim , Jae-Hyung Jeon

We observe the dynamics of waves propagating on the surface of a ferrofluid under the influence of a spatially and temporarily modulated field. In particular, we excite plane waves by a travelling lamellar modulation of the magnetization.…

Other Condensed Matter · Physics 2009-11-13 A. Beetz , C. Gollwitzer , R. Richter , I. Rehberg

We study wave propagation in networks of coupled cells which can behave as excitable or self-oscillatory media. For excitable media, an asymptotic construction of wave trains is presented. This construction predicts their shape and speed,…

Other Condensed Matter · Physics 2009-11-11 A. Carpio

Flagellated bacteria on nutrient-rich substrates can differentiate into a swarming state and move in dense swarms across surfaces. A recent experiment measured the flow in the fluid around an Escherichia coli swarm (Wu, Hosu and Berg, 2011…

Biological Physics · Physics 2016-07-28 Justas Dauparas , Eric Lauga

We show using theory and experiments that a small particle moving along an elastic membrane through a viscous fluid is repelled from the membrane due to hydro-elastic forces. The viscous stress field produces an elastic disturbance leading…

Soft Condensed Matter · Physics 2018-02-23 Bhargav Rallabandi , Naomi Oppenheimer , Matan Yah Ben Zion , Howard A. Stone

It is now well established that nearby beating pairs of eukaryotic flagella or cilia typically synchronize in phase. A substantial body of evidence supports the hypothesis that hydrodynamic coupling between the active filaments, combined…

Soft Condensed Matter · Physics 2016-10-12 Raymond E. Goldstein , Eric Lauga , Adriana I. Pesci , Michael R. E. Proctor
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