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The interaction between neighboring cracks has been shown to strongly influence the fracture behavior of graphene. While previous studies focused primarily on crack spacing, the role of crack width remains poorly understood. Here,…

Materials Science · Physics 2026-05-20 Suyeong Jin , Jung-Wuk Hong , Alexandre F. Fonseca

A concise method for following the evolving geometry of a moving surface using Lagrangian coordinates is described. All computations can be done in the fixed geometry of the initial surface despite the evolving complexity of the moving…

Soft Condensed Matter · Physics 2013-05-29 Mark A. Peterson

We investigate wrinkling of two-dimensional random and triangular semiflexible polymer networks under shear. Both types of semiflexible networks exhibit wrinkling above a small critical shear angle, which scales with an exponent of the…

Soft Condensed Matter · Physics 2014-06-06 Pascal Müller , Jan Kierfeld

"Necks" are features of lipid membranes characterized by an uniquley large curvature, functioning as bridges between different compartments. These features are ubiquitous in the life-cycle of the cell and instrumental in processes such as…

Soft Condensed Matter · Physics 2021-01-05 Piermarco Fonda , Luca Giomi

The eye grows during childhood to position the retina at the correct distance behind the lens to enable focused vision, a process called emmetropization. Animal studies have demonstrated that this growth process is dependent upon visual…

Biological Physics · Physics 2021-01-22 L. S. Kimpton , B. J. Walker , C. L. Hall , B. Bintu , D. Crosby , H. M. Byrne , A. Goriely

Accretion of mineralized thin wall-like structures via localized growth along their edges is observed in a range of physical and biological systems ranging from molluscan and brachiopod shells to carbonate-silica composite precipitates. To…

Soft Condensed Matter · Physics 2022-02-23 C. Nadir Kaplan , L. Mahadevan

A central question in developmental biology is how size and position are determined. The genetic code carries instructions on how to control these properties in order to regulate the pattern and morphology of structures in the developing…

Cell Behavior · Quantitative Biology 2016-12-05 Krishna Garikipati

When a droplet containing a concentrated suspension evaporates in a dry environment, a layer often forms at the interface accumulating non-volatile material. Such a "skin layer" experiences strong stresses and eventually turns mechanically…

Fluid Dynamics · Physics 2025-01-10 Raphael Saiseau , Lorenzo Botto , Christian Diddens , Alvaro Marin

Spatially confined rigid membranes reorganize their morphology in response to the imposed constraints. A crumpled elastic sheet presents a complex pattern of random folds focusing the deformation energy while compressing a membrane resting…

In comparative and developmental neuroanatomy one encounters questions regarding the deformation of neural tissue under stress. The motivation of this note is an observation (Barbas {\it et al}) that at cortical folds or gyri, the layers of…

Tissues and Organs · Quantitative Biology 2007-05-23 Partha P. Mitra

To deal with permanent deformations and residual stresses, we consider a morphoelastic model for the scar formation as the result of wound healing after a skin trauma. Next to the mechanical components such as strain and displacements, the…

Numerical Analysis · Mathematics 2022-03-17 Ginger Egberts , Fred Vermolen , Paul van Zuijlen

Growth-induced instabilities are ubiquitous in biological systems and lead to diverse morphologies in the form of wrinkling, folding, and creasing. The current work focusses on the mechanics behind growth-induced wrinkling instabilities in…

Pattern Formation and Solitons · Physics 2022-06-28 Sumit Mehta , Gangadharan Raju , Prashant Saxena

To go beyond the simple model for the fold as two flexible surfaces or faces linked by a crease that behaves as an elastic hinge, we carefully shape and anneal a crease within a polymer sheet and study its mechanical response. First, we…

Soft Condensed Matter · Physics 2018-08-16 Théo Jules , Frédéric Lechenault , Mokhtar Adda-Bedia

This review presents recent progress in understanding constraints and consequences of close-packing geometry of filamentous or columnar materials possessing non-trivial textures, focusing in particular on the common motifs of twisted and…

Soft Condensed Matter · Physics 2015-05-18 Gregory M. Grason

Inspired by recent experiments that highlight the role of nematic defects in the morphogenesis of epithelial tissues, we develop a minimal framework to study the dynamics of an active curved surface driven by its nematic texture. Allowing…

Soft Condensed Matter · Physics 2022-09-14 Farzan Vafa , L. Mahadevan

Transverse wrinkles are known to appear in thin rectangular elastic sheets when stretched in the long direction. Numerically computed bifurcation diagrams for extremely thin, highly stretched films indicate entire orbits of wrinkling…

Soft Condensed Matter · Physics 2025-12-02 Shrinidhi S. Pandurangi , Timothy J. Healey , Nicolas Triantafyllidis

The mechanical behaviour of solid biological tissues has long been described using models based on classical continuum mechanics. However, the classical continuum theories of elasticity and viscoelasticity cannot easily capture the…

Biological Physics · Physics 2017-05-16 Shakti N. Menon , Cameron L. Hall , Scott W. McCue , D. L. Sean McElwain

We present a newly developed approach for the calculation of interfacial stiffness and contact area evolution between two rough bodies exhibiting self affine surface structures. Using spline assisted discretization to define localised…

Soft Condensed Matter · Physics 2021-06-04 Dorian Hanaor , Yixiang Gan , Itai Einav

It is well known that an annular sheet could wrinkle as a result of axisymmetric tensile loads applied at the edges. In this system, regions under compression appear due to Poisson effect in the azimuthal direction yielding an incompatible…

Soft Condensed Matter · Physics 2019-06-19 I. Andrade-Silva , M. Adda-Bedia

A common developmental process, called branching morphogenesis, generates the epithelial trees in a variety of organs, including the lungs, kidneys, and glands. How branching morphogenesis can create epithelial architectures of very…

Tissues and Organs · Quantitative Biology 2021-03-02 Christine Lang , Lisa Conrad , Dagmar Iber