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The Sackin and Colless indices are two widely-used metrics for measuring the balance of trees and for testing evolutionary models in phylogenetics. This short paper contributes two results about the Sackin and Colless indices of trees. One…

Populations and Evolution · Quantitative Biology 2022-07-20 Gary Goh , Michael Fuchs , Louxin Zhang

Tree balance plays an important role in different research areas like theoretical computer science and mathematical phylogenetics. For example, it has long been known that under the Yule model, a pure birth process, imbalanced trees are…

Populations and Evolution · Quantitative Biology 2020-12-18 Mareike Fischer

Several indices that measure the degree of balance of a rooted phylogenetic tree have been proposed so far in the literature. In this work we define and study a new index of this kind, which we call the total cophenetic index: the sum, over…

Populations and Evolution · Quantitative Biology 2014-02-11 Arnau Mir , Francesc Rossello , Lucia Rotger

A phylogenetic network is a simplex (or 1-component tree-child) network if the child of every reticulation node is a network leaf. Simplex networks are a superclass of phylogenetic trees and a subclass of tree-child networks. Generalizing…

Populations and Evolution · Quantitative Biology 2022-01-03 Louxin Zhang

The study of species organization and their clustering by genetic or phenotypic similarity is carried out with the tools of phylogenetic trees. An important structural property of phylogenetic trees is the balance, which measures how taxa…

Populations and Evolution · Quantitative Biology 2020-09-01 T. Araújo Lima , Flavia M. D. Marquitti , Marcus A. M. de Aguiar

For two decades, the Colless index has been the most frequently used statistic for assessing the balance of phylogenetic trees. In this article, this statistic is studied under the Yule and uniform model of phylogenetic trees. The main tool…

Probability · Mathematics 2007-05-23 Michael G. B. Blum , Olivier François , Svante Janson

(Im)balance indices can be used to quantify the (im)balance of trees by assigning numerical scores to them. An easy way to generate a new index is to construct a compound index, e.g., a linear combination of established indices. Two of the…

Populations and Evolution · Quantitative Biology 2026-04-20 Linda Knüver , Mareike Fischer

In recent years, there has been an effort to extend the classical notion of phylogenetic balance, originally defined in the context of trees, to networks. One of the most natural ways to do this is with the so-called $B_2$ index. In this…

Populations and Evolution · Quantitative Biology 2024-07-30 François Bienvenu , Jean-Jil Duchamps , Michael Fuchs , Tsan-Cheng Yu

One of the main applications of balance indices is in tests of null models of evolutionary processes. The knowledge of an exact formula for a statistic of a balance index, holding for any number n of leaves, is necessary in order to use…

Populations and Evolution · Quantitative Biology 2012-10-22 Gabriel Cardona , Arnau Mir , Francesc Rossello

Measures of phylogenetic balance, such as the Colless and Sackin indices, play an important role in phylogenetics. Unfortunately, these indices are specifically designed for phylogenetic trees, and do not extend naturally to phylogenetic…

Populations and Evolution · Quantitative Biology 2022-11-14 François Bienvenu , Gabriel Cardona , Celine Scornavacca

Measures of tree balance play an important role in the analysis of phylogenetic trees. One of the oldest and most popular indices in this regard is the Colless index for rooted bifurcating trees, introduced by Colless (1982). While many of…

Populations and Evolution · Quantitative Biology 2020-02-18 Tomás M. Coronado , Mareike Fischer , Lina Herbst , Francesc Rosselló , Kristina Wicke

In mathematical phylogenetics, the time-consistent galled trees provide a simple class of rooted binary network structures that can be used to represent a variety of different biological phenomena. We study the enumerative combinatorics of…

Combinatorics · Mathematics 2025-04-24 Lily Agranat-Tamir , Michael Fuchs , Bernhard Gittenberger , Noah A. Rosenberg

We compute an explicit formula for the expected value of the Colless index of a phylogenetic tree generated under the Yule model, and an explicit formula for the expected value of the Sackin index of a phylogenetic tree generated under the…

Populations and Evolution · Quantitative Biology 2012-01-19 Arnau Mir , Francesc Rossello

Measures of tree balance play an important role in various research areas, for example in phylogenetics. There they are for instance used to test whether an observed phylogenetic tree differs significantly from a tree generated by the Yule…

Combinatorics · Mathematics 2019-07-23 Mareike Fischer , Lina Herbst , Kristina Wicke

Galled trees are studied as a recombination model in population genetics. This class of phylogenetic networks is generalized into tree-child, galled and reticulation-visible network classes by relaxing a structural condition imposed on…

Populations and Evolution · Quantitative Biology 2020-02-28 Gabriel Cardona , Louxin Zhang

Phylogenetic trees are important tools in the study of evolutionary relationships between species. Measures such as the index of Sackin, Colless, and Total Cophenetic have been extensively used to quantify tree balance, one key property of…

Populations and Evolution · Quantitative Biology 2020-09-01 T. Araújo Lima , Marcus A. M. de Aguiar

Measures of tree balance play an important role in many different research areas such as mathematical phylogenetics or theoretical computer science. Typically, tree balance is quantified by a single number which is assigned to the tree by a…

Populations and Evolution · Quantitative Biology 2025-07-31 Mareike Fischer , Tom Niklas Hamann , Kristina Wicke

Phylogenetic networks play an important role in evolutionary biology as, other than phylogenetic trees, they can be used to accommodate reticulate evolutionary events such as horizontal gene transfer and hybridization. Recent research has…

Populations and Evolution · Quantitative Biology 2024-03-19 Linda Knüver , Mareike Fischer , Marc Hellmuth , Kristina Wicke

The Colless index for bifurcating phylogenetic trees, introduced by Colless (1982), is defined as the sum, over all internal nodes $v$ of the tree, of the absolute value of the difference of the sizes of the clades defined by the children…

Populations and Evolution · Quantitative Biology 2020-07-30 Tomás M. Coronado , Arnau Mir , Francesc Rosselló

Tree balance plays an important role in phylogenetics and other research areas, which is why several indices to measure tree balance have been introduced over the years. Nevertheless, a formal definition of what a balance index actually is…

Populations and Evolution · Quantitative Biology 2023-11-10 Mareike Fischer , Lina Herbst , Sophie Kersting , Luise Kühn , Kristina Wicke
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