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Active particles, which interact hydrodynamically, display a remarkable variety of emergent collective phenomena. We use squirmers to model spherical microswimmers and explore the collective behavior of thousands of them under the influence…

Fluid Dynamics · Physics 2017-10-31 Jan-Timm Kuhr , Johannes Blaschke , Felix Rühle , Holger Stark

Geometric confinements are frequently encountered in soft matter systems and in particular significantly alter the dynamics of swimming microorganisms in viscous media. Surface-related effects on the motility of microswimmers can lead to…

Dense suspensions of swimming bacteria are known to exhibit collective behaviour arising from the interplay of steric and hydrodynamic interactions. Unconfined suspensions exhibit transient, recurring vortices and jets, whereas those…

Biological Physics · Physics 2016-07-04 Hugo Wioland , Enkeleida Lushi , Raymond E. Goldstein

Biological and artificial microswimmers often self-propel in external flows of vortical nature; relevant examples include algae in small-scale ocean eddies, spermatozoa in uterine peristaltic flows and bacteria in microfluidic devices. A…

Biological Physics · Physics 2022-11-14 Ivan Tanasijevic , Eric Lauga

Inspired by the classical Kepler and Rutherford problem, we investigate an analogous set-up in the context of active microswimmers: the behavior of a deformable microswimmer in a swirl flow. First we identify new steady bound states in the…

Soft Condensed Matter · Physics 2014-09-11 Mitsusuke Tarama , Andreas M. Menzel , Hartmut Löwen

Motivated by recent experiments of motile bacteria crossing liquid-liquid interfaces of isotropic- nematic coexistence (Cheon et al., Soft Matter 20: 7313-7320, 2024), we study the dynamics of prolate microswimmers traversing clean…

Soft Condensed Matter · Physics 2025-08-01 Rishish Mishra , Harish Pothukuchi , Harinadha Gidituri , Juho Lintuvuori

To explore and react to their environment, living micro-swimmers have developed sophisticated strategies for locomotion - in particular, motility with multiple gaits. To understand the physical principles associated with such a behavioural…

Swimming micro-organisms such as bacteria or spermatozoa are typically found in dense suspensions, and exhibit collective modes of locomotion qualitatively different from that displayed by isolated cells. In the dilute limit where…

Biological Physics · Physics 2010-05-02 Sebastien Michelin , Eric Lauga

We investigate the hydrodynamic interactions between microorganisms swimming at low Reynolds number. By considering simple model swimmers, and combining analytic and numerical approaches, we investigate the time-averaged flow field around a…

Soft Condensed Matter · Physics 2007-05-25 C. M. Pooley , G. P. Alexander , J. M. Yeomans

Studies of model microswimmers have significantly contributed to the understanding of the principles of self-propulsion we have today. However, only a small number of microswimmer types have been amenable to analytic modeling, and further…

Soft Condensed Matter · Physics 2021-02-10 Sebastian Ziegler , Maxime Hubert , Nicolas Vandewalle , Jens Harting , Ana-Sunčana-Smith

We address the problem of controlling a dynamical system governing the motion of a 3D weighted shape changing body swimming in a perfect fluid. The rigid displacement of the swimmer results from the exchange of momentum between prescribed…

Optimization and Control · Mathematics 2011-03-30 Thomas Chambrion , Alexandre Munnier

Magnetically driven artificial microswimmers have the potential to revolutionize many biomedical technologies, such as minimally-invasive microsurgery, micro-particle manipulation, and localized drug delivery. However, many of these…

Fluid Dynamics · Physics 2019-09-12 Jake Buzhardt , Phanindra Tallapragada

We study theoretically the behavior of a class of hydrodynamic dipoles. This study is motivated by recent experiments on synthetic and biological swimmers in microfluidic \textit{Hele-Shaw} type geometries. Under such confinement, a…

Fluid Dynamics · Physics 2015-09-30 Eva Kanso , Alan Tsang

The habitat of microorganisms is typically complex and viscoelastic. A natural question arises: Do polymers in a suspension of active swimmers enhance chaotic motion or promote orientational order? We address this issue by performing…

Soft Condensed Matter · Physics 2025-12-23 Yuan Zhou , Qingzhi Zou , Ignacio Pagonabarraga , Kaihuan Zhang , Kai Qi

Many microswimmers are inherently chiral, and this chirality can introduce fascinating behaviors in a collection of microswimmers. The dynamics become even more intriguing when two types of microswimmers with distinct chirality are mixed.…

Soft Condensed Matter · Physics 2025-09-03 Divya Kushwaha , Shradha Mishra

The question of characterization of the degree of non-equilibrium activity in active matter systems is studied in the context of a stochastic microswimmer model driven by a chemical cycle. The resulting dynamical properties and entropy…

Statistical Mechanics · Physics 2024-05-27 Michalis Chatzittofi , Jaime Agudo-Canalejo , Ramin Golestanian

Small organisms (e.g., bacteria) and artificial microswimmers move due to a combination of active swimming and passive Brownian motion. Considering a simplified linear three-sphere swimmer, we study how the swimmer size regulates the…

Soft Condensed Matter · Physics 2010-10-13 Jörn Dunkel , Irwin M. Zaid

In this letter we propose a kinematic model to show how collisions with a surface and rotational Brownian motion give rise to the accumulation of micro-swimmers near a surface. In this model, an elongated microswimmer invariably travels…

Biological Physics · Physics 2008-12-23 Guanglai Li , Jay X. Tang

We present the results of hydrodynamic simulations using the method of multi-particle collision dynamics for a system of squirmer microswimmers moving under the influence of gravity at low Reynolds numbers. In addition, the squirmers are…

Fluid Dynamics · Physics 2020-05-13 Felix Rühle , Holger Stark

We consider a suspension of active rigid particles (swimmers) in a steady Stokes flow, where particles are distributed according to a stationary ergodic random process, and we study its homogenization in the macroscopic limit. A key point…

Analysis of PDEs · Mathematics 2023-03-15 Armand Bernou , Mitia Duerinckx , Antoine Gloria
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