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We experimentally, numerically and analytically explore the diffusive transport of active colloidal particles with sensory delay, navigating motility landscapes in which the self-propulsion speed depends on space. We show how the transport…

Statistical Mechanics · Physics 2026-04-29 Adrià Garcés , Ueli Töpfer , Lucio Isa , Demian Levis , Ignacio Pagonabarraga

We introduce a system of bacteria confined to a finite 2D oil-water interface and driven on two distinct time scales by motility and by growth. The combined effect of activity on different time scales creates transitions between several…

Soft Condensed Matter · Physics 2023-06-28 Blake Langeslay , Gabriel Juarez

Collective motion of self-propelled organisms or synthetic particles often termed active fluid has attracted enormous attention in broad scientific community because of it fundamentally non-equilibrium nature. Energy input and interactions…

Soft Condensed Matter · Physics 2014-03-05 Shuang Zhou , Andrey Sokolov , Oleg D Lavrentovich , Igor S Aranson

Inspired by dense contractile tissues, where cells are subject to periodic deformation, we formulate and study a generic hydrodynamic theory of pulsating active liquids. Combining mechanical and phenomenological arguments, we postulate that…

Soft Condensed Matter · Physics 2025-09-25 Tirthankar Banerjee , Thibault Desaleux , Jonas Ranft , Étienne Fodor

A particle driven by an external force in a molecular crowding environment - a quiescent bath of other particles, makes their spatial distribution inhomogeneous: the bath particles accumulate in front of the biased particle (BP) and are…

Soft Condensed Matter · Physics 2011-09-08 Carlos Mejia-Monasterio , Gleb Oshanin

In equilibrium, the physical properties of matter are set by the interactions between the constituents. In contrast, the energy input of the individual components controls the behavior of synthetic or living active matter. Great progress…

Soft Condensed Matter · Physics 2025-08-29 Q. Martinet , Y. Li , A. Aubret , E. Hannezo , J. Palacci

Biological tissues have been observed to display emergent fluid-like properties, owing to physical interactions between cells. However, it remains unclear in general how these fluid-like properties affect tissue structure and function.…

Soft Condensed Matter · Physics 2026-02-24 Giulia L. Celora , Benjamin J. Walker , Mohit P. Dalwadi , Philip Pearce

Interactions between swimming cells and surfaces are essential to many microbiological processes, from bacterial biofilm formation to human fertilization. However, in spite of their fundamental importance, relatively little is known about…

Biological Physics · Physics 2013-02-04 Vasily Kantsler , Jörn Dunkel , Marco Polin , Raymond E. Goldstein

Microswimmers often exhibit surprising patterns due to the nonequilibrium nature of their dynamics. Collectively, suspensions of microswimmers appear as a liquid whose properties set it apart from its passive counterpart. To understand the…

Soft Condensed Matter · Physics 2019-05-22 Fabian Jan Schwarzendahl , Marco G. Mazza

The diffusion type is determined not only by microscopic dynamics but also by the environment properties. For example, the environment's fractal structure is responsible for the emergence of subdiffusive scaling of the mean square…

Statistical Mechanics · Physics 2021-10-26 Piotr Kubala , Michał Cieśla , Bartłomiej Dybiec

The movement of motor particles consisting of one or several molecular motors bound to a cargo particle is studied theoretically. The particles move on patterns of immobilized filaments. Several patterns are described for which the motor…

Statistical Mechanics · Physics 2009-11-11 Stefan Klumpp , Reinhard Lipowsky

Cells swimming in confined environments are attracted by surfaces. We measure the steady-state distribution of smooth-swimming bacteria (Escherichia coli) between two glass plates. In agreement with earlier studies, we find a strong…

Soft Condensed Matter · Physics 2008-10-02 Allison P. Berke , Linda Turner , Howard C. Berg , Eric Lauga

Active liquid crystals exert nonequilibrium stresses on their surroundings through constant consumption of energy, giving rise to dynamical steady states not present in equilibrium. The paradigmatic example of an active liquid crystal is a…

Soft Condensed Matter · Physics 2025-02-12 Paarth Gulati , Fernando Caballero , Itamar Kolvin , Zhihong You , M. Cristina Marchetti

The transport properties of colloidal particles in active liquids have been studied extensively. It has led to a deeper understanding of the interactions between passive and active particles. However, the phase behavior of colloidal…

Soft Condensed Matter · Physics 2023-01-30 Pragya Kushwaha , Vivek Semwal , Sayan Maity , Shraddha Mishra , Vijayakumar Chikkadi

Bacteria commonly live in structured communities that affect human health and influence ecological systems. Heterogeneous populations, such as motile and non-motile populations, often coexist in bacteria communities. Motile subpopulations…

Biological Physics · Physics 2019-06-19 Haoran Xu , Justas Dauparas , Debasish Das , Eric Lauga , Yilin Wu

Dense suspensions of swimming bacteria are known to exhibit collective behaviour arising from the interplay of steric and hydrodynamic interactions. Unconfined suspensions exhibit transient, recurring vortices and jets, whereas those…

Biological Physics · Physics 2016-07-04 Hugo Wioland , Enkeleida Lushi , Raymond E. Goldstein

Cells control fluid flows with a spatial and temporal precision that far exceeds the capabilities of current microfluidic technologies. Cells achieve this superior spatio-temporal control by harnessing dynamic networks of cytoskeleton and…

Soft Condensed Matter · Physics 2025-05-26 Fan Yang , Shichen Liu , Heun Jin Lee , Rob Phillips , Matt Thomson

Equilibrium self-assembly and conventional materials processing techniques fall far short of mimicking dynamic self-actuating processes that are commonplace throughout biology. To bridge the gap between living and synthetic matter, we study…

Ratchet effects can arise for single or collectively interacting Brownian particles on an asymmetric substrate when a net dc transport is produced by an externally applied ac driving force or by periodically flashing the substrate.…

Soft Condensed Matter · Physics 2017-12-06 C. J. Olson Reichhardt , C. Reichhardt

Swimmers and self-propelled particles are physical models for the collective behaviour and motility of a wide variety of living systems, such as bacteria colonies, bird flocks and fish schools. Such artificial active materials are amenable…