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Many animal cells change their shape depending on the stiffness of the substrate on which they are cultured: they assume small, rounded shapes in soft ECMs, they elongate within stiffer ECMs, and flatten out on hard substrates. Cells tend…

Cell Behavior · Quantitative Biology 2019-06-24 Elisabeth G. Rens , Roeland M. H. Merks

We introduce a novel model, comprising self-avoiding surfaces and incorporating edges and tubules, that is designed to characterize the structural morphologies and transitions observed within the endoplasmic reticulum (ER). By employing…

Soft Condensed Matter · Physics 2024-04-09 Jaya Kumar Alageshan , Yashodhan Hatwalne , Rahul Pandit

Cross-sections of cell shapes in a tissue monolayer typically resemble a tiling of convex polygons. Yet, examples exist where the polygons are not convex with curved cell-cell interfaces, as seen in the adaxial epidermis. To date,…

Biological Physics · Physics 2024-06-21 Kyungeun Kim , J. M. Schwarz , Martine Ben Amar

Most biological tissues grow by the synthesis of new material close to the tissue's interface, where spatial interactions can exert strong geometric influences on the local rate of growth. These geometric influences may be mechanistic, or…

Numerical Analysis · Mathematics 2020-05-28 Mohd Almie Alias , Pascal R Buenzli

We derive a model for the optimization of the bending and torsional rigidities of non-homogeneous elastic rods. This is achieved by studying a sharp interface shape optimization problem with perimeter penalization, that treats both…

Optimization and Control · Mathematics 2024-05-01 Patrick Dondl , Alberto Maione , Steve Wolff-Vorbeck

We address the folding induced by differential growth in soft layered solids via an elementary model that consists of a soft growing neo-Hookean elastic layer adhered to a deep elastic substrate. As the layer/substrate modulus ratio is…

Soft Condensed Matter · Physics 2015-09-30 Tuomas Tallinen , John S. Biggins

Animal morphogenesis involves complex tissue deformation processes, which require tight control over tissue rheology. Yet, it remains insufficiently understood how tissue rheology results from the interplay between cellular packing and…

The mechanical environment of a substrate plays a key role in influencing the behavior of adherent biological cells. Traditional tunable substrates have limitations as their mechanical properties cannot be dynamically altered in-situ during…

Soft Condensed Matter · Physics 2025-11-12 Manu Sebastian , Sreenath Balakrishnan , Safvan Palathingal

Rod-like bacteria maintain their cylindrical shapes with remarkable precision during growth. However, they are also capable to adapt their shapes to external forces and constraints, for example by growing into narrow or curved confinements.…

Subcellular Processes · Quantitative Biology 2016-02-10 Ariel Amir , Sven van Teeffelen

The majority of solid tumours arise in epithelia and therefore much research effort has gone into investigating the growth, renewal and regulation of these tissues. Here we review different mathematical and computational approaches that…

Tissues and Organs · Quantitative Biology 2019-12-23 O. J. Maclaren , A. G. Fletcher , H. M. Byrne , P. K. Maini

A new class of micromechanically motivated chain network models for soft biological tissues is presented. On the microlevel, it is based on the statistics of long chain molecules. A wormlike chain model is applied to capture the behavior of…

Quantitative Methods · Quantitative Biology 2009-11-10 E. Kuhl , K. Garikipati , E. M. Arruda , K. Grosh

The hydrodynamic theory of active nematics has been often used to describe the spatio-temporal dynamics of cell flows and motile topological defects within soft confluent tissues. Those theories, however, often rely on the assumption that…

Soft Condensed Matter · Physics 2023-08-15 Ioannis Hadjifrangiskou , Liam J. Ruske , Julia M. Yeomans

Collective cell migration governs a range of physiological and pathological processes, from tissue morphogenesis to cancer invasion, in which topological defects arise as an inevitable consequence of frequent cellular rearrangement and…

Soft Condensed Matter · Physics 2026-04-01 Jiusi Zhang , Chung Wing Chan , Bo Li , Rui Zhang

In the context of elasticity theory, rigidity theorems allow to derive global properties of a deformation from local ones. This paper presents a new asymptotic version of rigidity, applicable to elastic bodies with sufficiently stiff…

Analysis of PDEs · Mathematics 2019-09-04 Fabian Christowiak , Carolin Kreisbeck

We consider a free boundary problem for a system of PDEs, modeling the growth of a biological tissue. A morphogen, controlling volume growth, is produced by specific cells and then diffused and absorbed throughout the domain. The geometric…

Analysis of PDEs · Mathematics 2017-11-22 Alberto Bressan , Marta Lewicka

Shape transformations of epithelial tissues in three dimensions, which are crucial for embryonic development or in vitro organoid growth, can result from active forces generated within the cytoskeleton of the epithelial cells. How the…

Biological Physics · Physics 2024-12-23 Diana Khoromskaia , Guillaume Salbreux

There is increasing interest in the analysis of biological tissue, its organization and its dynamics with the help of mathematical models. In the ideal case emergent properties on the tissue scale can be derived from the cellular scale.…

Tissues and Organs · Quantitative Biology 2009-11-13 Tilo Beyer , Michael Meyer-Hermann

A constitutive relation between stress and strain relative to a reference state is the basic assumption of elasticity theory. However, in living matter, force generation is governed by motor molecule activity, which does not depend on…

Soft Condensed Matter · Physics 2026-03-17 Nikolas H. Claussen , Fridtjof Brauns , Boris I. Shraiman

We introduce an Active Vertex Model (AVM) for cell-resolution studies of the mechanics of confluent epithelial tissues consisting of tens of thousands of cells, with a level of detail inaccessible to similar methods. The AVM combines the…

Biological Physics · Physics 2016-12-20 Daniel L. Barton , Silke Henkes , Cornelis J. Weijer , Rastko Sknepnek

Over the last two decades, scientific literature has been blooming with various means of simulating epithelial cell colonies. Each of these simulations can be separated by their respective efficiency (expressed in terms of consumed…

Computational Physics · Physics 2022-12-20 Kevin Höllring , Ana-Sunčana Smith