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Pattern dynamics on curved surfaces are found everywhere in nature. The geometry of surfaces have been shown to influence dynamics and play a functional role, yet a comprehensive understanding is still elusive. Here, we report for the first…

Pattern Formation and Solitons · Physics 2022-06-15 Ryosuke Nishide , Shuji Ishihara

Turing's mechanism is often invoked to explain periodic patterns in nature, although direct experimental support is scarce. Turing patterns form in reaction-diffusion systems when the activating species diffuse much slower than the…

Biological Physics · Physics 2024-03-15 Lucas Menou , Chengjie Luo , David Zwicker

Reaction-diffusion (Turing) systems are fundamental to the formation of spatial patterns in nature and engineering. These systems are governed by a set of non-linear partial differential equations containing parameters that determine the…

Machine Learning · Computer Science 2022-11-28 Jordon Kho , Winston Koh , Jian Cheng Wong , Pao-Hsiung Chiu , Chin Chun Ooi

We consider pattern formation using reaction-diffusion equation on various non-uniformly curved surfaces. We explore how, in general, curvature and, in particular the domain shape would affect the pattern formation in these geometries. As…

Soft Condensed Matter · Physics 2017-05-08 Sankaran Nampoothiri , Amal Medhi

One possible mechanism to explain how animals got their coat patterns was proposed by Alan Turing. He assumed that two kinds of morphogens diffuse on a surface and interact with each other, generating a reaction-diffusion mechanism. We…

Tissues and Organs · Quantitative Biology 2017-11-16 Michäel Dougoud , Christian Mazza , Beat Schwaller , Laszlo Pecze

In certain biological contexts, such as the plumage patterns of birds and stripes on certain species of fishes, pattern formation takes place behind a so-called "wave of competency". Currently, the effects of a wave of competency on the…

Quantitative Methods · Quantitative Biology 2022-06-15 Yue Liu , Philip K. Maini , Ruth E. Baker

Cross-diffusion systems play a central role in mathematical modelling, in which density-dependent dispersal and multiscale mechanisms can lead to spatial segregation and diffusion-driven instabilities. In several relevant examples,…

Analysis of PDEs · Mathematics 2026-03-24 Brocchieri Elisabetta , Soresina Cinzia

Reaction-diffusion processes across layered media arise in several scientific domains such as pattern-forming E. coli on agar substrates, epidermal-mesenchymal coupling in development, and symmetry-breaking in cell polarisation. We develop…

Pattern Formation and Solitons · Physics 2020-09-18 Andrew L. Krause , Václav Klika , Jacob Halatek , Paul K. Grant , Thomas E. Woolley , Neil Dalchau , Eamonn A. Gaffney

We investigate Turing instability and pattern formation in two-dimensional domains for two reaction-diffusion models, obtained as diffusive limits of kinetic equations for mixtures of monatomic and polyatomic gases. The first model is of…

Mathematical Physics · Physics 2026-02-23 Stefano Boccelli , Giorgio Martalò , Romina Travaglini

We consider a two dimensional Turing like system with two diffusing species which interact with each other. Considering the species to be charged, we include the effect of an electric field along a given direction which can lead to a drift…

Other Condensed Matter · Physics 2008-12-31 B K Agarwalla , J K Bhattacharjee , P Titum

Turing instabilities for a two species reaction-diffusion systems is studied under anisotropic diffusion. More specifically, the diffusion constants which characterize the ability of the species to relocate in space are direction sensitive.…

Statistical Mechanics · Physics 2015-09-30 Daniel M. Busiello , Gwendoline Planchon , Malbor Asllani , Timoteo Carletti , Duccio Fanelli

Turing (or double-diffusive) instabilities describe pattern formation in reaction-diffusion systems, and were proposed in 1952 as a potential mechanism behind pattern formation in nature, such as leopard spots and zebra stripes. Because the…

Materials Science · Physics 2020-04-29 M. W. Noble , M. R. Tonks , S. P. Fitzgerald

Most of the physically based techniques for rendering translucent objects use the diffusion theory of light scattering in turbid media. The widely used dipole diffusion model (Jensen et al. 2001) applies the diffusion-theory formula derived…

Graphics · Computer Science 2010-11-16 Konstantin Kolchin

Turing theory of pattern formation is among the most popular theoretical means to account for the variety of spatio-temporal structures observed in Nature and, for this reason, finds applications in many different fields. While Turing…

Pattern Formation and Solitons · Physics 2025-10-22 Riccardo Muolo , Luca Gallo , Vito Latora , Mattia Frasca , Timoteo Carletti

Turing patterns are a central paradigm for describing spatial patterns in nature. The corresponding theory of reaction-diffusion dynamics combines ideal diffusion with nonlinear reactions, resulting in patterns when species diffuse at…

Biological Physics · Physics 2026-01-28 Cathelijne ter Burg , David Zwicker

The aim of this paper is to contribute to the understanding of the pattern formation phenomenon in reaction-diffusion equations coupled with ordinary differential equations. Such systems of equations arise, for example, from modeling of…

Analysis of PDEs · Mathematics 2016-07-15 Anna Marciniak-Czochra , Grzegorz Karch , Kanako Suzuki

We numerically solve the active nematohydrodynamic equations of motion, coupled to a Turing reaction-diffusion model, to study the effect of active nematic flow on the stripe patterns resulting from a Turing instability. If the activity is…

Soft Condensed Matter · Physics 2021-11-04 Saraswat Bhattacharyya , Julia M. Yeomans

From the striped coats of zebras to the ripples in windblown sand, the natural world abounds with locally banded patterns. Such patterns have been of great interest throughout history, and, in the last twenty years, scientists in a wide…

comp-gas · Physics 2016-08-31 David A. Egolf , Ilarion V. Melnikov , Eberhard Bodenschatz

Diffusion-driven patterns appear on curved surfaces in many settings, initiated by unstable modes of an underlying Laplacian operator. On a flat surface or perfect sphere, the patterns are degenerate, reflecting translational/rotational…

Soft Condensed Matter · Physics 2025-09-09 John R. Frank , Jemal Guven , Mehran Kardar , Leyna Shackleton

Reaction-diffusion waves in multiple spatial dimensions advance at a rate that strongly depends on the curvature of the wave fronts. These waves have important applications in many physical, ecological, and biological systems. In this work,…

Pattern Formation and Solitons · Physics 2022-12-28 Pascal R. Buenzli , Matthew J. Simpson
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