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In their search for metabolic resources microbes swim through viscous environments that present physical anisotropies, including steric obstacles across a wide range of sizes. Hydrodynamic forces are known to significantly alter swimmer…

Soft Condensed Matter · Physics 2020-07-03 Kentaro Hoeger , Tristan Ursell

Transport of material across liquid interfaces is ubiquitous for living cells and is also a crucial step in drug delivery and in many industrial processes. The fluids that are present on either side of the interfaces will usually have…

Soft Condensed Matter · Physics 2023-05-24 Chao Feng , John J. Molina , Matthew S. Turner , Ryoichi Yamamoto

Many biological fluids are composed of suspended polymers immersed in a viscous fluid. A prime example is mucus, where the polymers are also known to form a network. While the presence of this microstructure is linked with an overall…

Fluid Dynamics · Physics 2024-10-10 Adam K. Townsend , Eric E. Keaveny

We experimentally and theoretically study the dynamics of a low-Reynolds number helical swimmer moving across viscosity gradients. Experimentally, a double-layer viscosity is generated by superposing two miscible fluids with similar…

External gradients can strongly influence the collective behavior of microswimmers. In this paper, we study the behavior of two hydrodynamically interacting self-propelled chiral swimmers, in the low-Reynolds number regime, under the…

Biological Physics · Physics 2023-01-04 Ruma Maity , P. S. Burada

Swimming, i.e., being able to advance in the absence of external forces by performing cyclic shape changes, is particularly demanding at low Reynolds numbers which is the regime of interest for micro-organisms and micro-robots. We focus on…

Optimization and Control · Mathematics 2012-10-04 François Alouges , Laetitia Giraldi

Three-dimensional simulations with fully resolved hydrodynamics are performed to study the dynamics of a single squirmer under gravity, in order to clarify its motion in the vicinity of a flat plate. Different dynamics emerge for different…

Fluid Dynamics · Physics 2020-05-27 Federico Fadda , John Jairo Molina , Ryoichi Yamamoto

We study the orientational and translational dynamics of spherical microswimmers trapped at fluid interfaces, in terms of the force dipole and source dipole components of their flow field. Using numerical simulations and analytical…

Soft Condensed Matter · Physics 2022-04-26 Harinadha Gidituri , Zaiyi Shen , Alois Wurger , Juho S. Lintuvuori

Microscale propulsion is integral to numerous biomedical systems, for example biofilm formation and human reproduction, where the surrounding fluids comprise suspensions of polymers. These polymers endow the fluid with non-Newtonian…

Fluid Dynamics · Physics 2018-06-28 David A. Gagnon , Thomas D. Montenegro-Johnson

Active suspensions encompass a wide range of complex fluids containing microscale energy-injecting particles, such as cells, bacteria or artificially powered active colloids. Because they are intrinsically non-equilibrium, active…

Soft Condensed Matter · Physics 2024-07-09 Zhouyang Ge , Gwynn J. Elfring

The dynamics of a triangular magnetocapillary swimmer is studied using the lattice Boltzmann method. Performing extensive numerical simulations taking into account the coupled dynamics of the fluid-fluid interface and of magnetic particles…

Collective behaviour in suspensions of microswimmers is often dominated by the impact of long-ranged hydrodynamic interactions. These phenomena include active turbulence, where suspensions of pusher bacteria at sufficient densities exhibit…

Soft Condensed Matter · Physics 2019-08-27 Dora Bardfalvy , Henrik Nordanger , Cesare Nardini , Alexander Morozov , Joakim Stenhammar

Achieving macroscopic directed migration of microscale swimmers in a fluid is an important step towards utilizing their autonomous motion. It has been experimentally shown that directed motion can be induced, without any external fields, by…

Soft Condensed Matter · Physics 2017-09-14 Jiajun Tong , Michael J. Shelley

We consider the phase transition induced by compressing a self-avoiding walk in a slab where the walk is attached to both walls of the slab in two and three dimensions, and the resulting phase once the polymer is compressed. The process of…

Statistical Mechanics · Physics 2025-11-19 C J Bradly , N R Beaton , A L Owczarek

We study the dynamics of gyrotactic microswimmers suspended in homogeneous and isotropic turbulence by using direct numerical simulations (DNS). The swimmers are characterized by three non-dimensional parameters: their aspect ratio…

Soft Condensed Matter · Physics 2026-04-24 Suraj Kumar Nayak , Vishwanath Shukla , Akshay Bhatnagar

In this Letter, we study the collective behaviour of a large number of self-propelled microswimmers immersed in a fluid. Using unprecedently large-scale lattice Boltzmann simulations, we reproduce the transition to bacterial turbulence. We…

Soft Condensed Matter · Physics 2017-07-17 Joakim Stenhammar , Cesare Nardini , Rupert W. Nash , Davide Marenduzzo , Alexander Morozov

Suspensions of swimming micro-organisms are known to undergo intricate collective dynamics as a result of hydrodynamic and collision interactions. Micro-swimmers, such as bacteria and micro-algae, naturally live and have evolved in complex…

Fluid Dynamics · Physics 2025-10-31 Yasser Almoteri , Enkeleida Lushi

We study the effects of hydrodynamic interactions between a wall and the Purcell three-link swimmer in the two-dimensional case. After deriving the equations of motion in a low Reynolds number regime using Resistive Force Theory with…

Fluid Dynamics · Physics 2026-03-19 Enrico Micalizio , Marco Morandotti , Henry Shum , Marta Zoppello

We address the problem of controlling a dynamical system governing the motion of a 3D weighted shape changing body swimming in a perfect fluid. The rigid displacement of the swimmer results from the exchange of momentum between prescribed…

Optimization and Control · Mathematics 2011-03-30 Thomas Chambrion , Alexandre Munnier

Living organisms employ diverse strategies to navigate confined environments. Inspired by translocation observations on California blackworms (\textit{Lumbriculus variegatus}), we combine biological experiments and active-polymer…

Soft Condensed Matter · Physics 2025-10-21 K. R. Prathyusha , Paulami Sarkar , Justin Xu , Saad Bhamla