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Symmetry breaking is essential for biological microswimmers to achieve locomotion in viscous environments. Such asymmetry in the swimming mechanism enables the generation of directional forces that overcome fluid resistance, leading to…

Fluid Dynamics · Physics 2025-12-29 Anupriya Dutta Roy , Smita S. Sontakke , Arvind Kumar , Ranabir Dey , Anupam Gupta

Suspensions of swimming micro-organisms are known to undergo intricate collective dynamics as a result of hydrodynamic and collision interactions. Micro-swimmers, such as bacteria and micro-algae, naturally live and have evolved in complex…

Fluid Dynamics · Physics 2025-10-31 Yasser Almoteri , Enkeleida Lushi

The dynamic interactions between pairs of swimming microorganisms underpin the collective behaviour of larger suspensions, but accurately calculating pairwise collisions has typically required the use of numerical simulations in which…

Soft Condensed Matter · Physics 2021-11-30 C. Darveniza , T. Ishikawa , T. J. Pedley , D. R. Brumley

We study the fluid drift due to a time-dependent dumbbell model of a microswimmer. The model captures important aspects of real microswimmers such as a time-dependent flagellar motion and a no-slip body. The model consists of a rigid sphere…

Soft Condensed Matter · Physics 2017-02-01 Peter Mueller , Jean-Luc Thiffeault

In this review we discuss the recent progress in the simulation of soft active matter systems and in particular the hydrodynamics of microswimmers using the method of multiparticle collision dynamics, which solves the hydrodynamic flows…

Soft Condensed Matter · Physics 2020-09-03 Andreas Zöttl

The time dynamics of flagellar and ciliary beating is often neglected in theories of microswimmers, with the most common models prescribing a time-constant actuation of the surrounding fluid. By explicitly introducing a metachronal wave,…

Soft Condensed Matter · Physics 2026-01-19 G. C. Antunes , H. Stark

The behaviour of microscopic swimmers has previously been explored near large scale confining geometries and in the presence of very small-scale surface roughness. Here we consider an intermediate case of how a simple microswimmer, the…

Fluid Dynamics · Physics 2021-09-14 Kenta Ishimoto , Eamonn A. Gaffney , David J. Smith

In this note, we study the effect of viscosity gradients on the energy dissipated by the motion of microswimmers and the associated efficiency of that motion. Using spheroidal squirmer model swimmers in weak linearly varying viscosity…

Fluid Dynamics · Physics 2024-11-20 Jiahao Gong , Vaseem A. Shaik , Gwynn J. Elfring

We theoretically study the self-propulsion of a thin (slender) colloid driven by asymmetric chemical reactions on its surface at vanishing Reynolds number. Using the method of matched asymptotic expansions, we obtain the colloid…

Soft Condensed Matter · Physics 2018-03-21 Yahaya Ibrahim , Ramin Golestanian , Tanniemola B. Liverpool

Ciliated microswimmers and flagellated bacteria alter their swimming trajectories to follow the direction of an applied electric field exhibiting electrotaxis. Both for matters of application and physical modelling, it is instructive to…

Soft Condensed Matter · Physics 2024-01-26 Carola M. Buness , Avi Rana , Corinna C. Maass , Ranabir Dey

Many aquatic microorganisms are able to swim. In natural environments they typically do so in the presence of flows. In recent years it has been shown that the interplay of swimming and flows can give rise to interesting and biologically…

Fluid Dynamics · Physics 2022-07-20 M. Borgnino , G. Boffetta , M. Cencini , F. De Lillo , K. Gustavsson

Self-propelled particles move along circles rather than along a straight line when their driving force does not coincide with their propagation direction. Examples include confined bacteria and spermatozoa, catalytically driven nanorods,…

Soft Condensed Matter · Physics 2008-08-18 Sven van Teeffelen , Hartmut Löwen

Several micro-organisms, such as bacteria, algae, or spermatozoa, use flagella or cilia to swim in a fluid, while many other micro-organisms instead use ample shape deformation, described as amoeboid, to propel themselves by either crawling…

Biological Physics · Physics 2016-09-19 Hao Wu , A. Farutin , W. -F. Hu , M. Thiébaud , S. Rafaï , P. Peyla , M. -C. Lai , C. Misbah

Many micro-swimmers propel themselves by rotating micro-cylindrical organelles such as flagella or cilia. These cylindrical organelles almost never live in free space, yet their motions in a confining geometry can be counter-intuitive. For…

Fluid Dynamics · Physics 2023-12-25 Hanliang Guo , Yi Man , Hai Zhu

External gradients can strongly influence the collective behavior of microswimmers. In this paper, we study the behavior of two hydrodynamically interacting self-propelled chiral swimmers, in the low-Reynolds number regime, under the…

Biological Physics · Physics 2023-01-04 Ruma Maity , P. S. Burada

We derive from first principles a three-dimensional theory of self-propelled particle swarming in a viscous fluid environment. Our model predicts emergent collective behavior that depends critically on fluid opacity, mechanism of…

Soft Condensed Matter · Physics 2026-05-12 Yao-Li Chuang , M. R. D'Orsogna , T. Chou

In isotropic fluids like water, micrometer-scale swimmers have evolved swim strokes to translate despite their tiny size. As described by Purcell in his Scallop Theorem, reciprocal motions, like those performed by a scallop, cannot drive…

Locomotion is typically studied either in continuous media where bodies and legs experience forces generated by the flowing medium, or on solid substrates dominated by friction. In the former, centralized coordination is believed to…

Biological Physics · Physics 2023-03-29 Baxi Chong , Juntao He , Shengkai Li , Eva Erickson , Kelimar Diaz , Tianyu Wang , Daniel Soto , Daniel I. Goldman

The paper presents a two-phase hydrodynamic model for the numerical simulation of collective motion in a thin layer of active colloids containing spherical microswimmers. The model accounts for three fundamental mechanisms governing the…

Soft Condensed Matter · Physics 2025-12-10 A. Kiverin , S. Luguev , I. Yakovenko

Microorganisms are rarely found in Nature swimming freely in an unbounded fluid. Instead, they typically encounter other organisms, hard walls, or deformable boundaries such as free interfaces or membranes. Hydrodynamic interactions between…

Fluid Dynamics · Physics 2013-10-21 Marcelo A. Dias , Thomas R. Powers