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Related papers: Metachronal coordination at the mesoscale

200 papers

It has long been known that some microswimmers seem to swim counter-intuitively faster when the viscosity of the surrounding fluid is increased, whereas others slow down. This conflicting dependence of the swimming velocity on the viscosity…

Soft Condensed Matter · Physics 2016-11-08 Jayant Pande , Laura Merchant , Timm Krüger , Jens Harting , Ana-Sunčana Smith

We define a model microswimmer with a variable cycle time, thus allowing the possibility of phase locking driven by hydrodynamic interactions between swimmers. We find that, for extensile or contractile swimmers, phase locking does occur,…

Soft Condensed Matter · Physics 2015-05-13 Victor B. Putz , Julia M. Yeomans

We examine the hydrodynamic performance of two cilia beating patterns reconstructed from experimental data. In their respective natural systems, the two beating patterns correspond to: (A) pumping-specialized cilia, and (B)…

Fluid Dynamics · Physics 2015-06-19 Hanliang Guo , Janna Nawroth , Yang Ding , Eva Kanso

Microswimming cells and robots exhibit diverse behaviours due to both their swimming and their environment. One of the core environmental features impacting inertialess swimming is background flows. While the influence of select flows,…

Fluid Dynamics · Physics 2025-10-30 Eamonn A. Gaffney , Kenta Ishimoto , Benjamin J. Walker

Metamaterials are structures composed of repeating unit-cells which enable macro-scale properties not found in nature. Since metamaterials are typically solid structures with predetermined interconnections, it is challenging to leverage…

Fluid Dynamics · Physics 2025-12-10 Ezra Ben-Abu , Anna Zigelman , Sefi Givli , Amir D. Gat

On surfaces with many motile cilia, beats of the individual cilia coordinate to form metachronal waves. We present a theoretical framework that connects the dynamics of an individual cilium to the collective dynamics of a ciliary carpet via…

Soft Condensed Matter · Physics 2021-03-12 Fanlong Meng , Rachel R. Bennett , Nariya Uchida , Ramin Golestanian

The time dynamics of flagellar and ciliary beating is often neglected in theories of microswimmers, with the most common models prescribing a time-constant actuation of the surrounding fluid. By explicitly introducing a metachronal wave,…

Soft Condensed Matter · Physics 2026-01-19 G. C. Antunes , H. Stark

Self-propelled colloids (swimmers) in confining geometries follow trajectories determined by hydrodynamic interactions with the bounding surfaces. However, typically these interactions are ignored or truncated to lowest order. We…

We investigate synchronization caused by long-range hydrodynamic interaction in a two-dimensional, substrated array of rotors with random intrinsic frequencies. The rotor mimics a flagellated bacterium that is attached to the substrate…

Statistical Mechanics · Physics 2010-03-30 Nariya Uchida , Ramin Golestanian

We introduce a measure for energetic efficiency of biological cilia acting individually or collectively and numerically determine the optimal beating patterns according to this criterion. Maximizing the efficiency of a single cilium leads…

Biological Physics · Physics 2011-09-21 Natan Osterman , Andrej Vilfan

We study synchronization in bulk suspensions of spherical microswimmers with chiral trajectories using large scale numerics. The model is generic. It corresponds to the lowest order solution of a general model for self-propulsion at low…

Soft Condensed Matter · Physics 2023-01-18 Sotiris Samatas , Juho S. Lintuvuori

Metamaterials are artificial composite structures designed for controlling waves or fields, and exhibit interaction phenomena that are unexpected on the basis of their chemical constituents. These phenomena are encoded in effective material…

Materials Science · Physics 2015-12-21 Mihai Caleap , Bruce W Drinkwater

Motile cilia beat in an asymmetric fashion in order to propel the surrounding fluid. When many cilia are located on a surface, their beating can synchronise such that their phases form metachronal waves. Here, we computationally study a…

Soft Condensed Matter · Physics 2023-09-29 David J. Hickey , Ramin Golestanian , Andrej Vilfan

Studies of model microswimmers have significantly contributed to the understanding of the principles of self-propulsion we have today. However, only a small number of microswimmer types have been amenable to analytic modeling, and further…

Soft Condensed Matter · Physics 2021-02-10 Sebastian Ziegler , Maxime Hubert , Nicolas Vandewalle , Jens Harting , Ana-Sunčana-Smith

Synchronization induced by long-range hydrodynamic interactions is attracting attention as a candidate mechanism behind coordinated beating of cilia and flagella. Here we consider a minimal model of hydrodynamic synchronization in the low…

Soft Condensed Matter · Physics 2012-11-26 Nariya Uchida , Ramin Golestanian

We suggest several reciprocal swimming mechanisms that lead to a locomotion only in viscoelastic fluids. The first situation is to have a difference between the two amplitudes of the oscillatory arm motion for a three-sphere microswimmer.…

Soft Condensed Matter · Physics 2021-06-08 Kento Yasuda , Mizuki Kuroda , Shigeyuki Komura

Hydrodynamic and magnetohydrodynamic numerical studies of a mechanically forced two-vortex flow inside a sphere are reported. The simulations are performed in the intermediate regime between the laminar flow and developed turbulence where a…

Fluid Dynamics · Physics 2015-05-14 K. Reuter , F. Jenko , A. Tilgner , C. B. Forest

Many active biological particles, such as swimming microorganisms or motor-proteins, do work on their environment by going though a periodic sequence of shapes. Interactions between particles can lead to the phase-synchronization of their…

Soft Condensed Matter · Physics 2022-07-26 Brato Chakrabarti , Michael J. Shelley , Sebastian Fürthauer

This study introduces a novel method to break wave transmission reciprocity by leveraging the synchronization of self-oscillators. An experimental demonstration with aeroacoustic cavities is presented. They behave as weakly nonlinear limit…

Applied Physics · Physics 2025-08-25 Alexander K. Stoychev , Ulrich Kuhl , Nicolas Noiray

A variety of swimming microorganisms, called ciliates, exploit the bending of a large number of small and densely-packed organelles, termed cilia, in order to propel themselves in a viscous fluid. We consider a spherical envelope model for…

Fluid Dynamics · Physics 2011-08-30 Sebastien Michelin , Eric Lauga