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Biological microswimmers often encounter deformable boundaries in physiological conditions; for instance, the viscoelastic walls of reproductive tract during migration of spermatozoa, or host tissue during early bacterial biofilm formation.…

Soft Condensed Matter · Physics 2025-08-07 Smita S. Sontakke , Aneesha Kajampady , Mohd Suhail Rizvi , Ranabir Dey

Microorganisms ofter move in confined, disordered environments, where hydrodynamic couplings can modify their transport behavior. Using extensive finite-element simulations, we investigate the dynamics of microswimmers -- modeled as…

Soft Condensed Matter · Physics 2026-03-24 Mirko Residori , Sebastian Aland , Christina Kurzthaler

Bacteria in biofilms form complex structures and can collectively migrate within mobile aggregates, which is referred to as swarming. This behavior is influenced by a combination of various factors, including morphological characteristics…

Soft Condensed Matter · Physics 2024-01-19 Bohan Wu-Zhang , Dmitry A. Fedosov , Gerhard Gompper

Active matter exhibits various forms of non-equilibrium states in the absence of external forcing, including macroscopic steady-state currents. Such states are often too complex to be modelled from first principles and our understanding of…

Soft Condensed Matter · Physics 2020-09-17 Viktor Škultéty , Cesare Nardini , Joakim Stenhammar , Davide Marenduzzo , Alexander Morozov

Efficient locomotion is important for the evolution of complex life, yet the physical principles selecting specific swimming strokes often remain entangled with biological constraints. In viscous fluids, the scallop theorem constrains the…

Biological Physics · Physics 2026-03-11 Takahiro Kanazawa , Kenta Ishimoto , Kyogo Kawaguchi

The swimming of a spheroid immersed in a viscous fluid and performing surface deformations periodically in time is studied on the basis of Stokes equations of low Reynolds number hydrodynamics. The average over a period of time of the…

Fluid Dynamics · Physics 2016-11-23 B. U. Felderhof

Self-propelled microparticles create flow fields that determine how they interact with surfaces, external flows, and each other. These flow fields fall into distinct classes--pushers, pullers, and neutral swimmers--each exhibiting…

Soft Condensed Matter · Physics 2025-12-25 Lisa Rohde , Gordei Anchutkin , Viktor Holubec , Frank Cichos

We study the dynamics of gyrotactic microswimmers suspended in homogeneous and isotropic turbulence by using direct numerical simulations (DNS). The swimmers are characterized by three non-dimensional parameters: their aspect ratio…

Soft Condensed Matter · Physics 2026-04-24 Suraj Kumar Nayak , Vishwanath Shukla , Akshay Bhatnagar

We study the coupled hydrodynamics between a motile slender microswimmer and a deformable interface that separates two Newtonian fluid regions. From the disturbance field generated by the swimming motion, we quantitatively characterize the…

Soft Condensed Matter · Physics 2022-12-22 Sankalp Nambiar , John S Wettlaufer

Hydrodynamic interaction strongly influences the collective behavior of the microswimmers. With this work, we study the behavior of two hydrodynamically interacting self-propelled chiral swimmers in the low Reynolds number regime,…

Soft Condensed Matter · Physics 2022-11-28 Ruma Maity , P. S. Burada

We study numerically the hydrodynamics of a self-propelled particle system, consisting of spherical squirmers sedimented on a flat surface. We observe the emergence of dynamic structures, due to the interplay of particle-particle and…

Soft Condensed Matter · Physics 2019-12-20 Zaiyi Shen , Juho S. Lintuvuori

We calculate the hydrodynamic force on a small spherical, unsteady squirmer moving with a time-dependent velocity in a fluid at rest, taking into account convective and unsteady fluid inertia effects in perturbation theory. Our results…

Fluid Dynamics · Physics 2023-12-20 T. Redaelli , F. Candelier , R. Mehaddi , C. Eloy , B. Mehlig

The dynamics of sedimenting particles under gravity are surprisingly complex due to the presence of effective long-ranged forces. When the particles are polar with a well-defined symmetry axis and non-uniform density, recent theoretical…

Soft Condensed Matter · Physics 2023-02-22 Kavinda J. Nissanka , Xiaolei Ma , Justin C. Burton

We examine the dynamics of small anisotropic particles (spheroids) sedimenting through homogeneous isotropic turbulence using direct numerical simulations and theory. The gravity-induced inertial torque acting on sub-Kolmogorov spheroids…

Atmospheric and Oceanic Physics · Physics 2020-07-22 Prateek Anand , Samriddhi Sankar Ray , Ganesh Subramanian

We analyse a simple 'Stokesian squirmer' model for the enhanced mixing due to swimming micro-organisms. The model is based on a calculation of Thiffeault & Childress [Physics Letters A, 374, 3487 (2010), arXiv:0911.5511], where fluid…

Fluid Dynamics · Physics 2013-09-24 Zhi Lin , Jean-Luc Thiffeault , Stephen Childress

We investigate how shear influences the dynamics of active particles in dilute to concentrated suspensions. Using apolar active suspensions of squirmers as model systems, we show how their long-time diffusive dynamics can surprisingly slow…

Soft Condensed Matter · Physics 2025-11-14 Zhouyang Ge , John F. Brady , Gwynn J. Elfring

Microswimmers and active colloids often move in confined systems, including those involving interfaces. Such interfaces, especially at the microscale, may deform in response to the stresses of the flow created by the active particle. We…

Soft Condensed Matter · Physics 2026-01-14 Adam Hitin Bialus , Bhargav Rallabandi , Naomi Oppenheimer

We investigate the orientation dynamics of a neutrally buoyant spheroid, of an arbitrary aspect ratio ($\kappa$), freely rotating in a weakly viscoelastic fluid undergoing simple shear flow. Weak elasticity is characterized by a small but…

Fluid Dynamics · Physics 2025-05-30 Pavan Kumar Singeetham , Deepak Madival , Piyush Garg , Ganesh Subramanian

We study the dynamics of a prolate spheroidal microswimmer in Poiseuille flow for different flow geometries. When moving between two parallel plates or in a cylindrical microchannel, the swimmer performs either periodic swinging or periodic…

Soft Condensed Matter · Physics 2013-01-24 Andreas Zöttl , Holger Stark

When swimming animals form cohesive groups, they can reap several benefits. Our understanding of collective animal motion has traditionally been driven by models based on phenomenological behavioral rules, but more recent work has…

Fluid Dynamics · Physics 2024-11-15 Mohamed Niged Mabrouk , Daniel Floryan