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Self-propulsion allows living systems to display unusual collective behavior. Unlike passive systems in thermal equilibrium, active matter systems are not constrained by conventional thermodynamic laws. A question arises however as to what…

Soft Condensed Matter · Physics 2015-03-18 Sho C. Takatori , John F. Brady

Swimmers and self-propelled particles are physical models for the collective behaviour and motility of a wide variety of living systems, such as bacteria colonies, bird flocks and fish schools. Such artificial active materials are amenable…

We study an inertial chiral active fluid, formed by repulsive particles that transfer angular momentum through odd interactions, i.e. transverse forces. Chirality induces an inhomogeneous phase, consisting of rotating bubbles, whose…

Soft Condensed Matter · Physics 2026-05-06 Alessandro Petrini , Raphaël Maire , Umberto Marini Bettolo Marconi , Lorenzo Caprini

Active transport of biomolecular condensates and cell migration in collectives are fundamental to development, homeostasis, and processes such as cancer progression, wound healing, and infection response. Yet how these assemblies are…

Soft Condensed Matter · Physics 2025-10-24 Hossein Vahid , Jens-Uwe Sommer , Abhinav Sharma

Motility induced phase separation is an efficient aggregation mechanism of active matter, yet biological systems exhibit richer organization through communication among constituents. We investigate suspensions of active particles that…

Soft Condensed Matter · Physics 2025-08-18 Yuxin Zhou , Qingqing Yin , Shubhadip Nayak , Poulami Bag , Pulak K. Ghosh , Yunyun Li , Fabio Marchesoni

Semi-flexible filaments in living systems are constantly driven by active forces that often organize into mesoscale coherent flows. Although theory and simulations predict rich filament dynamics, experimental studies of passive filaments in…

Soft Condensed Matter · Physics 2026-01-09 Bipul Biswas , Devadyouti Das , Manasa Kandula , Shuang Zhou

The general problem of tracer diffusion in non-equilibrium baths is important in a wide range of systems, from the cellular level to geographical lengthscales. In this paper, we revisit the archetypical example of such a system: a…

Soft Condensed Matter · Physics 2023-11-08 Henrik Nordanger , Alexander Morozov , Joakim Stenhammar

Ratchet effects can arise for single or collectively interacting Brownian particles on an asymmetric substrate when a net dc transport is produced by an externally applied ac driving force or by periodically flashing the substrate.…

Soft Condensed Matter · Physics 2017-12-06 C. J. Olson Reichhardt , C. Reichhardt

Solvent plays an important role in the relative motion of nanoscopic bodies, and the study of such phenomena can help elucidate the mechanism of hydrophobic assembly, as well as the influence of solvent-mediated effects on in vivo motion in…

Statistical Mechanics · Physics 2011-12-12 Joseph A. Morrone , Jingyuan Li , B. J. Berne

We study interacting active Brownian particles (ABPs) with a space-dependent swim velocity via simulation and theory. We find that, although an equation of state exists, a mechanical equilibrium does not apply to ABPs in activity…

Soft Condensed Matter · Physics 2024-12-13 Adam Wysocki , Anil K. Dasanna , Heiko Rieger

Thanks to a constant energy input, active matter can self-assemble into phases with complex architectures and functionalities such as living clusters that dynamically form, reshape and break-up, which are forbidden in equilibrium materials…

Soft Condensed Matter · Physics 2019-03-27 Falko Schmidt , Benno Liebchen , Hartmut Löwen , Giovanni Volpe

We experimentally determine the force exerted by a bath of active particles onto a passive probe as a function of its distance to a wall and compare it to the measured averaged density distribution of active particles around the probe.…

Soft Condensed Matter · Physics 2022-08-10 Shuvojit Paul , Ashreya Jayaram , N Narinder , Thomas Speck , Clemens Bechinger

While the behavior of active colloidal molecules is well studied by now for a constant activity, the effect of activity gradients is much less understood. Here we explore one of the simplest molecules in activity gradients, namely active…

Soft Condensed Matter · Physics 2022-10-19 Pietro Luigi Muzzeddu , Hidde Derk Vuijk , Hartmut Lowen , Jens-Uwe Sommer , Abhinav Sharma

The nonequilibrium structural and dynamical properties of semiflexible active polar polymers subject to linear flow are studied by numerical simulations. Filaments are confined in two dimensions and immersed in a fluid described by the…

Soft Condensed Matter · Physics 2026-04-23 A. Lamura , R. G. Winkler

Some nonequilibrium systems exhibit anomalous suppression of the large-scale density fluctuations, so-called hyperuniformity. Recently, hyperuniformity was found numerically in a simple model of chiral active fluids [Q.-L. Lei et al., Sci.…

Soft Condensed Matter · Physics 2023-11-13 Yuta Kuroda , Kunimasa Miyazaki

We introduce for the first time a general model of biased-active particles, where the direction of the active force has a biased angle from the principle orientation of the anisotropic interaction between particles. We find that a highly…

Soft Condensed Matter · Physics 2018-06-07 Yongliang Gou , Huijun Jiang , Zhoughuai Hou

We study the emergent orientational dynamics of a dumbbell dimer -- two asymmetric monomers connected by a linking spring -- in a three-dimensional chiral environment with odd viscosity. In classical systems with conserved parity symmetry,…

Soft Condensed Matter · Physics 2025-10-23 Michalis Chatzittofi , Yuto Hosaka

The diffusive behavior of small entities is strongly influenced by the flow of the surrounding medium, which is ubiquitous in natural and artificial environments. In this study, we investigate the transport characteristics of the inertial…

Soft Condensed Matter · Physics 2025-11-19 Ankit Gupta , P. S. Burada

Assembly of protein complexes like virus shells, the centriole, the nuclear pore complex or the actin cytoskeleton is strongly determined by their spatial structure. Moreover it is becoming increasingly clear that the reversible nature of…

Subcellular Processes · Quantitative Biology 2014-05-13 Heinrich C. R. Klein , Ulrich S. Schwarz

We investigate the emergent interactions between two active Brownian particles coupled by an attractive harmonic potential and in contact with a thermal reservoir. By analyzing the stationary distribution of their separation, we demonstrate…

Statistical Mechanics · Physics 2025-10-08 Ritwick Sarkar , Sreya Chatterjee , Urna Basu