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Large Language Models (LLMs) are composed of neurons that exhibit various behaviors and roles, which become increasingly diversified as models scale. Recent studies have revealed that not all neurons are active across different datasets,…

Computation and Language · Computer Science 2024-03-19 Haoyun Xu , Runzhe Zhan , Derek F. Wong , Lidia S. Chao

Identifying important neurons for final predictions is essential for understanding the mechanisms of large language models. Due to computational constraints, current attribution techniques struggle to operate at neuron level. In this paper,…

Computation and Language · Computer Science 2024-09-26 Zeping Yu , Sophia Ananiadou

While a lot of analysis has been carried to demonstrate linguistic knowledge captured by the representations learned within deep NLP models, very little attention has been paid towards individual neurons.We carry outa neuron-level analysis…

Computation and Language · Computer Science 2020-10-07 Nadir Durrani , Hassan Sajjad , Fahim Dalvi , Yonatan Belinkov

Despite their widespread use, the mechanisms by which large language models (LLMs) represent and regulate uncertainty in next-token predictions remain largely unexplored. This study investigates two critical components believed to influence…

Machine Learning · Computer Science 2024-11-11 Alessandro Stolfo , Ben Wu , Wes Gurnee , Yonatan Belinkov , Xingyi Song , Mrinmaya Sachan , Neel Nanda

Large language models struggle with representing and generating rare tokens despite their importance in specialized domains. In this study, we identify neuron structures with exceptionally strong influence on language model's prediction of…

Artificial Intelligence · Computer Science 2025-05-23 Jing Liu , Haozheng Wang , Yueheng Li

Language models have been shown to perform better with an increase in scale on a wide variety of tasks via the in-context learning paradigm. In this paper, we investigate the hypothesis that the ability of a large language model to…

Computation and Language · Computer Science 2023-08-17 Hritik Bansal , Karthik Gopalakrishnan , Saket Dingliwal , Sravan Bodapati , Katrin Kirchhoff , Dan Roth

In recent years, the rapid advancement of large language models (LLMs) in natural language processing has sparked significant interest among researchers to understand their mechanisms and functional characteristics. Although prior studies…

Neurons and Cognition · Quantitative Biology 2026-01-08 Yiheng Liu , Zhengliang Liu , Zihao Wu , Junhao Ning , Haiyang Sun , Sichen Xia , Yang Yang , Xiaohui Gao , Ning Qiang , Bao Ge , Tianming Liu , Junwei Han , Xintao Hu

Large language models (LLMs) display strong comprehensive abilities, yet the internal mechanisms that support these behaviors remain insufficiently understood. In this work, we show that across a wide range of open-weight Transformers, a…

Machine Learning · Computer Science 2026-05-29 Xiangtian Ji , Yuxin Chen , Zhengzhou Cai , Xiang Wang , An Zhang , Tat-Seng Chua

Despite rapid adoption and deployment of large language models (LLMs), the internal computations of these models remain opaque and poorly understood. In this work, we seek to understand how high-level human-interpretable features are…

Machine Learning · Computer Science 2023-06-06 Wes Gurnee , Neel Nanda , Matthew Pauly , Katherine Harvey , Dmitrii Troitskii , Dimitris Bertsimas

As large language models (LLMs) continue to advance, their capacity to function effectively across a diverse range of languages has shown marked improvement. Preliminary studies observe that the hidden activations of LLMs often resemble…

Computation and Language · Computer Science 2025-06-12 Yuxin Chen , Yiran Zhao , Yang Zhang , An Zhang , Kenji Kawaguchi , Shafiq Joty , Junnan Li , Tat-Seng Chua , Michael Qizhe Shieh , Wenxuan Zhang

Recent developments in transformer-based language models have allowed them to capture a wide variety of world knowledge that can be adapted to downstream tasks with limited resources. However, what pieces of information are understood in…

Computation and Language · Computer Science 2024-01-31 Shrayani Mondal , Rishabh Garodia , Arbaaz Qureshi , Taesung Lee , Youngja Park

Understanding and shaping the behaviour of Large Language Models (LLMs) is increasingly important as applications become more powerful and more frequently adopted. This paper introduces a machine unlearning method specifically designed for…

Machine Learning · Computer Science 2024-07-25 Nicholas Pochinkov , Nandi Schoots

Biological neurons exhibit remarkable intelligence: they maintain internal states, communicate selectively with other neurons, and self-organize into complex graphs rather than rigid hierarchical layers. What if artificial intelligence…

Machine Learning · Computer Science 2025-12-01 Antoine Salomon

While a lot of work has been done in understanding representations learned within deep NLP models and what knowledge they capture, little attention has been paid towards individual neurons. We present a technique called as Linguistic…

Computation and Language · Computer Science 2024-01-17 Nadir Durrani , Fahim Dalvi , Hassan Sajjad

Large language models (LLMs) have revolutionized the field of natural language processing (NLP), and recent studies have aimed to understand their underlying mechanisms. However, most of this research is conducted within a monolingual…

Computation and Language · Computer Science 2025-09-29 Weixuan Wang , Barry Haddow , Minghao Wu , Wei Peng , Alexandra Birch

Advances in Large Language Models (LLMs) have led to remarkable capabilities, yet their inner mechanisms remain largely unknown. To understand these models, we need to unravel the functions of individual neurons and their contribution to…

Machine Learning · Computer Science 2023-06-01 Alex Foote , Neel Nanda , Esben Kran , Ioannis Konstas , Shay Cohen , Fazl Barez

Understanding the function of individual neurons within language models is essential for mechanistic interpretability research. We propose $\textbf{Neuron to Graph (N2G)}$, a tool which takes a neuron and its dataset examples, and…

Machine Learning · Computer Science 2023-04-26 Alex Foote , Neel Nanda , Esben Kran , Ionnis Konstas , Fazl Barez

Large Language Models (LLMs) typically have billions of parameters and are thus often difficult to interpret in their operation. In this work, we demonstrate that it is possible to decode neuron weights directly into token probabilities…

Computation and Language · Computer Science 2025-03-03 Harry J Davies

Self-Organized Operational Neural Networks (Self-ONNs) have recently been proposed as new-generation neural network models with nonlinear learning units, i.e., the generative neurons that yield an elegant level of diversity; however, like…

Computer Vision and Pattern Recognition · Computer Science 2023-04-18 Serkan Kiranyaz , Junaid Malik , Mehmet Yamac , Mert Duman , Ilke Adalioglu , Esin Guldogan , Turker Ince , Moncef Gabbouj

Probing large language models (LLMs) has yielded valuable insights into their internal mechanisms by linking neural activations to interpretable semantics. However, the complex mechanisms that link neuron's functional co-activation with the…

Computation and Language · Computer Science 2026-01-30 Yu Zheng , Yuan Yuan , Yue Zhuo , Yong Li , Gabriel Kreiman , Tomaso Poggio , Paolo Santi
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