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Microswimmers in nature often experience spatial gradients of viscosity. In this work we develop theoretical results for the dynamics of active particles, biological or otherwise, swimming through viscosity gradients. We model the active…

Fluid Dynamics · Physics 2019-10-16 Charu Datt , Gwynn J. Elfring

Active particles (living or synthetic) often move through inhomogeneous environments, such as gradients in light, heat or nutrient concentration, that can lead to directed motion (or taxis). Recent research has explored inhomogeneity in the…

Soft Condensed Matter · Physics 2022-08-23 Jiahao Gong , Vaseem A. Shaik , Gwynn J. Elfring

Organisms often swim through fluids that are spatially inhomogeneous. If the fluids are polymeric, gradients in polymer concentration may lead to gradients in both fluid viscosity and elasticity. In this letter, we present theoretical…

Fluid Dynamics · Physics 2024-11-08 Vaseem A. Shaik , Jiahao Gong , Gwynn J. Elfring

In this paper, we explore the hydrodynamics of spheroidal active particles in viscosity gradients. This work provides a more accurate modeling approach, in comparison to spherical particles, for anisotropic organisms like Paramecium…

Fluid Dynamics · Physics 2025-07-09 Jiahao Gong , Vaseem A. Shaik , Gwynn J. Elfring

Active particles, which interact hydrodynamically, display a remarkable variety of emergent collective phenomena. We use squirmers to model spherical microswimmers and explore the collective behavior of thousands of them under the influence…

Fluid Dynamics · Physics 2017-10-31 Jan-Timm Kuhr , Johannes Blaschke , Felix Rühle , Holger Stark

In this work, we analyze the motion of an active particle, modeled as a spherical squirmer, in linearly varying viscosity fields. In general, the presence of a particle will disturb a background viscosity field and the disturbance generated…

Fluid Dynamics · Physics 2021-07-09 Vaseem A. Shaik , Gwynn J. Elfring

Many fascinating properties of biological active matter crucially depend on the capacity of constituting entities to perform directed motion, e.g., molecular motors transporting vesicles inside cells or bacteria searching for food. While…

Statistical Mechanics · Physics 2023-06-21 Pietro Luigi Muzzeddu , Édgar Roldán , Andrea Gambassi , Abhinav Sharma

Experiments have shown that self-propelled particles can slide along the surface of a circular obstacle without becoming trapped over long times. Using simulations and theory, we study the impact of boundary conditions on the diffusive…

Biological Physics · Physics 2019-01-30 Theresa Jakuszeit , Ottavio A. Croze , Samuel Bell

The experiments of Leptos et al. [Phys. Rev. Lett. 103, 198103 (2009)] show that the displacements of small particles affected by swimming microorganisms achieve a non-Gaussian distribution, which nevertheless scales diffusively -- the…

Soft Condensed Matter · Physics 2017-10-26 Jean-Luc Thiffeault

For natural microswimmers, the interplay of swimming activity and external flow can promote robust motion, e.g. propulsion against ("upstream rheotaxis") or perpendicular to the direction of flow. These effects are generally attributed to…

Soft Condensed Matter · Physics 2018-01-30 Jaideep Katuri , William E. Uspal , Juliane Simmchen , Albert Miguel-López , Samuel Sánchez

We investigate theoretically the collective dynamics of soft active particles living in a viscous fluid. We focus on a minimal model for active but non-motile particles consisting of $N>1$ elastic dimers deformed by active stresses and…

Soft Condensed Matter · Physics 2010-04-09 Denis Bartolo , Eric Lauga

We study the behaviour of interacting self-propelled particles, whose self-propulsion speed decreases with their local density. By combining direct simulations of the microscopic model with an analysis of the hydrodynamic equations obtained…

Statistical Mechanics · Physics 2012-06-25 F. D. C. Farrell , J. Tailleur , D. Marenduzzo , M. C. Marchetti

We show that dense active fluids comprising interacting particles with persistent self-propulsion are driven to a non-equilibrium steady state consisting of co-moving particles with co-aligned active forces. This velocity and force sorting…

Soft Condensed Matter · Physics 2025-09-03 Suman Dutta , Pinaki Chaudhuri , Madan Rao , Chandan Dasgupta

Micron-sized self-propelled (active) particles can be considered as model systems for characterizing more complex biological organisms like swimming bacteria or motile cells. We produce asymmetric microswimmers by soft lithography and study…

Particles such as microplastics and phytoplankton suspended in the water column in the natural environment are often subject to the action of surface gravity waves. By modelling such anisotropic particles as small spheroids that slowly…

Fluid Dynamics · Physics 2023-08-03 Nimish Pujara , Jean-Luc Thiffeault

Most classical work on the hydrodynamics of low-Reynolds-number swimming addresses deterministic locomotion in quiescent environments. Thermal fluctuations in fluids are known to lead to a Brownian loss of the swimming direction. As most…

Fluid Dynamics · Physics 2014-06-18 Mario Sandoval , Navaneeth K. M. , Ganesh Subramanian , Eric Lauga

The effect of crowding on the run-and-tumble dynamics of swimmers such as bacteria is studied using a discrete lattice model of mutually excluding particles that move with constant velocity along a direction that is randomized at a rate…

Soft Condensed Matter · Physics 2015-06-16 Rodrigo Soto , Ramin Golestanian

Microswimmers in suspension exhibit collective swimming behaviour, forming various self-organised structures including ordered, aggregated, and turbulent-like structures. When mixed with passive particles phase-separation is known to occur,…

Fluid Dynamics · Physics 2026-03-26 Alexander Chamolly , Takuji Ishikawa

We find the mixing induced by a small swimming organism in a density stratified fluid. We model the swimmer as a spherical squirmer and quantify mixing through a mixing efficiency that is the ratio of rate of change of potential energy of…

Soft Condensed Matter · Physics 2025-02-26 Vaseem A. Shaik , Gwynn J. Elfring

Many biological active agents respond to gradients of environmental cues by redirecting their motion. Besides the well-studied prominent examples such as photo- and chemotaxis, there has been considerable recent interest in topotaxis, i.e.\…

Soft Condensed Matter · Physics 2023-05-04 Zeinab Sadjadi , Heiko Rieger
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