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Bacterial swarms display intriguing dynamical states like active turbulence. Using a hydrodynamic model we now show that such dense active suspensions manifest super-diffusion, via L\'evy walks, which masquerades as a crossover from…

Soft Condensed Matter · Physics 2021-09-15 Siddhartha Mukherjee , Rahul K. Singh , Martin James , Samriddhi Sankar Ray

We investigate the effects of heterogeneous (spatially varying) activity in a hydrodynamical model for dense bacterial suspensions, confining ourselves to experimentally realizable, simple, quenched, activity patterns. We show that the…

Soft Condensed Matter · Physics 2026-05-12 Siddhartha Mukherjee , Kunal Kumar , Samriddhi Sankar Ray

Active particles such as swimming bacteria or self-propelled colloids are known to spontaneously organize into fascinating large-scale dynamic structures. The emergence of these collective states from the motility pattern of the individual…

Soft Condensed Matter · Physics 2019-11-20 Hamid Karan , Gerardo E. Pradillo , Petia M. Vlahovska

Recent experiments (G. Ariel, et al., Nature Comm. 6, 8396 (2015)) revealed an intriguing behavior of swarming bacteria: they fundamentally change their collective motion from simple diffusion into a superdiffusive L\'{e}vy walk dynamics.…

Statistical Mechanics · Physics 2017-04-05 Sergei Fedotov , Nickolay Korabel

Dense suspensions of swimming bacteria are known to exhibit collective behaviour arising from the interplay of steric and hydrodynamic interactions. Unconfined suspensions exhibit transient, recurring vortices and jets, whereas those…

Biological Physics · Physics 2016-07-04 Hugo Wioland , Enkeleida Lushi , Raymond E. Goldstein

We experimentally study the emergence of collective bacterial swimming, a phenomenon often referred to as bacterial turbulence. A phase diagram of the flow of 3D E. coli suspensions spanned by bacterial concentration, the swimming speed of…

Soft Condensed Matter · Physics 2021-03-09 Yi Peng , Zhengyang Liu , Xiang Cheng

Suspensions of swimming bacteria interact hydrodynamically over long ranges, organizing themselves into collective states that drive large-scale chaotic flows, often referred to as "bacterial turbulence". Despite extensive experimental and…

We show that Lagrangian measurements in active turbulence bear imprints of turbulent and anomalous streaky hydrodynamics leading to a self-selection of persistent trajectories - Levy walks - over diffusive ones. This emergent dynamical…

Fluid Dynamics · Physics 2022-08-26 Rahul K. Singh , Siddhartha Mukherjee , Samriddhi Sankar Ray

Bacterial assemblies exhibit rich collective behaviors that control their biological functions, making them a relevant object of study from an active matter physics perspective. Dense bacterial suspensions self-organize into distinct…

Soft Condensed Matter · Physics 2025-04-08 Martin Maliet , Nicolas Fix-Boulier , Ludovic Berthier , Maxime Deforet

Dense bacterial suspensions at fluid interfaces provide a natural platform to explore active turbulence in a dimensional mismatch: active units are restricted to a two-dimensional surface, while the induced flows extend into the surrounding…

Soft Condensed Matter · Physics 2025-11-18 Yuanfeng Yin , Bokai Zhang , H. P. Zhang , Shuo Guo

Turbulence is ubiquitous, from oceanic currents to small-scale biological and quantum systems. Self-sustained turbulent motion in microbial suspensions presents an intriguing example of collective dynamical behavior amongst the simplest…

New aspects of turbulence are uncovered if one considers flow motion from the perspective of a fluid particle (known as the Lagrangian approach) rather than in terms of a velocity field (the Eulerian viewpoint). Using a new experimental…

Fluid Dynamics · Physics 2009-11-07 N. Mordant , J. Delour , E. Leveque , A. Arneodo , J. -F. Pinton

The field of active matter explores the behaviors of self propelled agents out of equilibrium, with active suspensions, such as swimming bacteria in solutions, serving as impactful models. These systems exhibit spatio-temporal patterns akin…

Soft Condensed Matter · Physics 2025-08-26 Pratikshya Jena , Shradha Mishra

We investigate the effects of bacterial activity on the mixing and transport properties of a passive scalar in time-periodic flows in experiments and in a simple model. We focus on the interactions between swimming E. coli and the…

Fluid Dynamics · Physics 2024-06-07 Ranjiangshang Ran , Paulo E. Arratia

In this Letter, we study the collective behaviour of a large number of self-propelled microswimmers immersed in a fluid. Using unprecedently large-scale lattice Boltzmann simulations, we reproduce the transition to bacterial turbulence. We…

Soft Condensed Matter · Physics 2017-07-17 Joakim Stenhammar , Cesare Nardini , Rupert W. Nash , Davide Marenduzzo , Alexander Morozov

Self-sustained turbulent structures have been observed in a wide range of living fluids, yet no quantitative theory exists to explain their properties. We report experiments on active turbulence in highly concentrated 3D suspensions of…

Dense suspensions of self-propelled bacteria and related active fluids exhibit spontaneous flow generation, vortex formation, and spatiotemporally chaotic dynamics despite operating at vanishingly small Reynolds numbers. These phenomena,…

Soft Condensed Matter · Physics 2026-02-26 Sandip Sahoo , Siddhartha Mukherjee , Samriddhi Sankar Ray

Swimming microorganisms rely on effective mixing strategies to achieve efficient nutrient influx. Recent experiments, probing the mixing capability of unicellular biflagellates, revealed that passive tracer particles exhibit anomalous…

Soft Condensed Matter · Physics 2013-02-04 Irwin M. Zaid , Jörn Dunkel , Julia M. Yeomans

Self-organized dynamic patterns in dense active matter are striking manifestations of non-equilibrium physics. A prominent example is the macroscopic elliptical motion observed in quasi-2D bacterial suspensions, which has lacked a physical…

Soft Condensed Matter · Physics 2026-03-30 Bing Miao , Lei-Han Tang

Collective motion is a phenomenon observed across length scales in nature, from bacterial swarming and tissue migration to the flocking of animals. The mechanisms underlying this behavior vary significantly depending on the biological…

Soft Condensed Matter · Physics 2025-01-15 Urvi Mahendra Bora , Mohd Suhail Rizvi
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