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The trajectories of microswimmers moving in narrow channels of widths comparable to their sizes are significantly altered when they encounter another microswimmer moving in the opposite direction. The consequence of these encounters is a…

The spatiotemporal dynamics in systems of active self-propelled particles is controlled by the propulsion mechanism in combination with various direct interactions, such as steric repulsion, hydrodynamics, and chemical fields. Yet, these…

Soft Condensed Matter · Physics 2024-06-03 Mario Theers , Elmar Westphal , Kai Qi , Roland G. Winkler , Gerhard Gompper

An artificial microswimmer drifts in response to spatio-temporal modulations of an activating suspension medium. We consider two competing mechanisms capable of influencing its tactic response: angular fluctuations, which help it explore…

Soft Condensed Matter · Physics 2017-02-15 Alexander Geiseler , Peter Hänggi , Fabio Marchesoni

Swimming organisms in their natural habitat navigate through a wide array of geometries and chemical environments. Interaction with the boundaries is ubiquitous and can significantly modify the swimming characteristics of the organism as…

Fluid Dynamics · Physics 2010-12-23 Saikat Jana , Soong Ho Um , Sunghwan Jung

Cooperation is vital for the survival of a swarm$^1$. Large scale cooperation allows murmuring starlings to outmaneuver preying falcons$^2$, shoaling sardines to outsmart sea lions$^3$, and homo sapiens to outlive their Pleistocene…

Soft Condensed Matter · Physics 2021-01-01 Matan Yah Ben Zion , Alvin Modin , Yaelin Caba , Paul M Chaikin

In several biologically relevant situations, cell locomotion occurs in polymeric fluids with Weissenberg {number} larger than one. Here we present results of three-dimensional numerical simulations for the steady locomotion of a…

Fluid Dynamics · Physics 2012-12-03 Lailai ZHu , Minh Do-Quang , Eric Lauga , Luca Brandt

Typical bodily and environmental fluids encountered by biological swimmers consist of dissolved macromolecules such as proteins and polymers, often rendering them non Newtonian. To mimic such scenarios, we investigate the motion of swimming…

Soft Condensed Matter · Physics 2023-05-23 Prateek Dwivedi , Atishay Shrivastava , Dipin Pillai , Rahul Mangal

We study, by means of an exact analytical solution, the motion of a spheroidal, axisymmetric squirmer in an unbounded fluid, as well as the low Reynolds number hydrodynamic flow associated to it. In contrast to the case of a spherical…

Fluid Dynamics · Physics 2020-11-30 Ruben Poehnl , Mihail N. Popescu , William E. Uspal

Micro-organisms usually can swim in their liquid environment by flagellar or ciliary beating. In this numerical work, we analyze the influence of flagellar beating on the orbits of a swimming cell in a shear flow. We also calculate the…

Soft Condensed Matter · Physics 2017-11-21 Levan Jibuti , Walter Zimmermann , Salima Rafaï , Philippe Peyla

Micron-sized self-propelled (active) particles can be considered as model systems for characterizing more complex biological organisms like swimming bacteria or motile cells. We produce asymmetric microswimmers by soft lithography and study…

Biological microswimmers often encounter deformable boundaries in physiological conditions; for instance, the viscoelastic walls of reproductive tract during migration of spermatozoa, or host tissue during early bacterial biofilm formation.…

Soft Condensed Matter · Physics 2025-08-07 Smita S. Sontakke , Aneesha Kajampady , Mohd Suhail Rizvi , Ranabir Dey

We suggest several reciprocal swimming mechanisms that lead to a locomotion only in viscoelastic fluids. The first situation is to have a difference between the two amplitudes of the oscillatory arm motion for a three-sphere microswimmer.…

Soft Condensed Matter · Physics 2021-06-08 Kento Yasuda , Mizuki Kuroda , Shigeyuki Komura

Various aspects of self-motility of chemically active colloids in Newtonian fluids can be captured by simple models for their chemical activity plus a phoretic slip hydrodynamic boundary condition on their surface. For particles of simple…

Soft Condensed Matter · Physics 2018-04-09 M. N. Popescu , W. E. Uspal , Z. Eskandari , M. Tasinkevych , S. Dietrich

Unicellular microscopic organisms living in aqueous environments outnumber all other creatures on Earth. A large proportion of them are able to self-propel in fluids with a vast diversity of swimming gaits and motility patterns. In this…

Biological Physics · Physics 2021-07-14 Marcos F. Velho Rodrigues , Maciej Lisicki , Eric Lauga

Although the motility of the flagellated bacteria, Escherichia coli, has been widely studied, the effect of viscosity on swimming speed remains controversial. The swimming mode of wild-type E.coli is often idealized as a "run-and- tumble"…

Biological Physics · Physics 2018-05-09 Zijie Qu , Fatma Zeynep Temel , Rene Henderikx , Kenneth S. Breuer

Self-propulsion of cellular microswimmers generates flow signatures, commonly classified as pusher- and puller-type, which characterize hydrodynamic interactions with other cells or boundaries. Using experimentally measured beat patterns,…

Cell Behavior · Quantitative Biology 2016-01-20 Gary S. Klindt , Benjamin M. Friedrich

The dynamic interactions between pairs of swimming microorganisms underpin the collective behaviour of larger suspensions, but accurately calculating pairwise collisions has typically required the use of numerical simulations in which…

Soft Condensed Matter · Physics 2021-11-30 C. Darveniza , T. Ishikawa , T. J. Pedley , D. R. Brumley

We present a new development of the force-coupling method (FCM) to address the accurate simulation of a large number of interacting micro-swimmers. Our approach is based on the squirmer model, which we adapt to the FCM framework, resulting…

Soft Condensed Matter · Physics 2015-12-21 Blaise Delmotte , Eric Keaveny , Franck Plouraboue , Eric Climent

We investigate the hydrodynamic interactions between microorganisms swimming at low Reynolds number. By considering simple model swimmers, and combining analytic and numerical approaches, we investigate the time-averaged flow field around a…

Soft Condensed Matter · Physics 2007-05-25 C. M. Pooley , G. P. Alexander , J. M. Yeomans

Ensembles of biological and artificial microswimmers produce long-range velocity fields with strong nonequilibrium fluctuations, which result in dramatic increase diffusivity of embedded particles (tracers). While such enhanced diffusivity…

Soft Condensed Matter · Physics 2019-02-20 Sergey Belan , Mehran Kardar