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Dynamical mechanisms that can stabilize the coexistence or diversity in biology are generally of fundamental interest. In contrast to many two-strategy evolutionary games, games with three strategies and cyclic dominance like the…

Populations and Evolution · Quantitative Biology 2010-03-15 Markus Schütt , Jens Christian Claussen

We study the influence of a randomly switching reproduction-predation rate on the survival behavior of the non-spatial cyclic Lotka-Volterra model, also known as the zero-sum rock-paper-scissors game, used to metaphorically describe the…

Populations and Evolution · Quantitative Biology 2018-02-28 Robert West , Mauro Mobilia , Alastair M. Rucklidge

We study the phenomenon of cyclic dominance in the paradigmatic Rock--Paper--Scissors model, as occurring in both stochastic individual-based models of finite populations and in the deterministic replicator equations. The mean-field…

Populations and Evolution · Quantitative Biology 2017-08-03 Qian Yang , Tim Rogers , Jonathan H P Dawes

We study a three-species cyclic model whose organisms are vulnerable to contamination with an infectious disease which propagates person-to-person. We consider that individuals of one species perform an evolutionary self-preservation…

Populations and Evolution · Quantitative Biology 2024-06-05 J. Menezes

We investigate the population dynamics in generalized Rock-Paper-Scissors models with an arbitrary number of species $N$. We show, for the first time, that spiral patterns with $N$-arms may develop both for odd and even $N$, in particular…

Biological Physics · Physics 2012-10-16 P. P. Avelino , D. Bazeia , L. Losano , J. Menezes , B. F. Oliveira

Several theoretical models predict that spatial patterning increases ecosystem resilience. However, these predictions rely on simplifying assumptions, such as assuming isotropic and infinitely large ecosystems, and empirical evidence…

Populations and Evolution · Quantitative Biology 2026-04-06 David Pinto-Ramos , Ricardo Martinez-Garcia

Rock-scissors-paper game, as the simplest model of intransitive relation between competing agents, is a frequently quoted model to explain the stable diversity of competitors in the race of surviving. When increasing the number of…

Physics and Society · Physics 2019-05-21 D. Bazeia , B. F. de Oliveira , A. Szolnoki

We analyze the replicator-mutator equations for the Rock-Paper-Scissors game. Various graph-theoretic patterns of mutation are considered, ranging from a single unidirectional mutation pathway between two of the species, to global…

Dynamical Systems · Mathematics 2015-06-11 Danielle F. P. Toupo , Steven H. Strogatz

The spatial rock-scissors-paper game (or cyclic Lotka-Volterra system) is extended to study how the spatiotemporal patterns are affected by the constructed backgrounds providing uniform number of neighbors (degree) at each site. On the…

Statistical Mechanics · Physics 2009-11-10 Gyorgy Szabo , Attila Szolnoki , Rudolf Izsak

Ecosystems frequently display the coexistence of diverse species under resource competition, typically resulting in skewed distributions of rarity and abundance. A potential driver of such coexistence is environmental fluctuations that…

Populations and Evolution · Quantitative Biology 2025-08-07 Davide Zanchetta , Deepak Gupta , Sofia Moschin , Samir Suweis , Amos Maritan , Sandro Azaele

What determines species diversity is dramatic concern in science. Here we report the effect of doping on diversity in spatiotemporal rock-paper-scissors (RPS) games, which can be observed directly in ecological, biological and social…

Adaptation and Self-Organizing Systems · Physics 2007-10-11 Wang Zhijian

There is mounting evidence that species interactions often involve long-term memory, with highly-varying waiting times between successive events and long-range temporal correlations. Accounting for memory undermines the common Markovian…

Populations and Evolution · Quantitative Biology 2025-11-24 Ohad Vilk , Mauro Mobilia , Michael Assaf

Predicting species persistence within ecological communities is a fundamental challenge for both empirical and theoretical ecology. Existing methods span from mechanistic models, whose parameters are difficult to estimate from data, to…

Populations and Evolution · Quantitative Biology 2026-04-30 Davide Bernardi , Giorgio Nicoletti , Prajwal Padmanabha , Samir Suweis , Sandro Azaele , Simon A. Levin , Andrea Rinaldo , Amos Maritan

Cyclic dominance of competing species is an intensively used working hypothesis to explain biodiversity in certain living systems, where the evolutionary selection principle would dictate a single victor otherwise. Technically the…

Populations and Evolution · Quantitative Biology 2021-06-16 D. Bazeia , M. J. B. Ferreira , B. F. de Oliveira , A. Szolnoki

Game dynamics in which three or more strategies are cyclically competitive, as represented by the rock-scissors-paper game, have attracted practical and theoretical interests. In evolutionary dynamics, cyclic competition results in…

Populations and Evolution · Quantitative Biology 2008-03-11 Naoki Masuda

Noise and spatial degrees of freedom characterize most ecosystems. Some aspects of their influence on the coevolution of populations with cyclic interspecies competition have been demonstrated in recent experiments [e.g. B. Kerr et al.,…

Populations and Evolution · Quantitative Biology 2007-12-08 Tobias Reichenbach , Mauro Mobilia , Erwin Frey

Competition between species and genotypes is a dominant factor in a variety of ecological and evolutionary processes. Biological dynamics are typically highly stochastic, and therefore, analyzing a competitive system requires accounting for…

Populations and Evolution · Quantitative Biology 2025-01-09 Ori Turkia , Nadav M. Shnerb

The composition of ecological communities varies not only between different locations but also in time. Understanding the fundamental processes that drive species towards rarity or abundance is crucial to assessing ecosystem resilience and…

Populations and Evolution · Quantitative Biology 2024-11-22 Emil Mallmin , Arne Traulsen , Silvia De Monte

Natural birth and death are fundamental mechanisms of population dynamics in ecosystems and have played pivotal roles in shaping population dynamics. Nevertheless, in studies of cyclic competition systems governed by the rock-paper-scissors…

Populations and Evolution · Quantitative Biology 2026-02-19 Yikang Lu , Wenhao She , Xiaofang Duan , Junpyo Park

We study diffusion on a multilayer network where the contact dynamics between the nodes is governed by a random process and where the waiting time distribution differs for edges from different layers. We study the impact on a random walk of…

Physics and Society · Physics 2018-12-18 Martin Gueuning , Sibo Cheng , Renaud Lambiotte , Jean-Charles Delvenne
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